2011 in arthropod paleontology

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This list of fossil arthropods described in 2012 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that have been described during the year 2012. The list only includes taxa at the level of genus or species.

Arachnids

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Acarus indicus^[1]	Sp. nov	Disputed	Kumar et al.	Early Permian	Manjir Formation	India	A mite. Originally described as a species of Acarus; Sidorchuk (2018) considered it more likely to be a misidentified immature nothrine oribatid mite (probably a member of the family Nothridae), and more likely to be a contaminant from the extant soil fauna rather than a genuine Paleozoic fossil.^[2]
Ameticos^[3]	Gen. et sp. nov		Garwood et al.	Carboniferous (late Stephanian)		France	A member of Opiliones. The type species is A. scolos.
Burmesiola^[4]	Gen. et sp. nov	Valid	Wunderlich	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A tree trunk spider. Genus includes new species B. cretacea.
Chaerilobuthus^[5]	Gen. et sp. nov	Valid	Lourenço & Beigel	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A scorpion. The type species is C. complexus.
Eoscaphiella^[4]	Gen. et sp. nov	Valid	Wunderlich	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Tetrablemmidae. Genus includes new species E. ohlhoffi.
Episinus tibiaseta^[6]	Sp. nov	Valid	Wunderlich	Miocene	Dominican amber	Dominican Republic	A member of the family Theridiidae, a species of Episinus.
Femurraptor^[6]	Gen. et sp. nov	Valid	Wunderlich	Miocene	Dominican amber	Dominican Republic	A member of the family Theridiidae. Genus includes new species F. dominicanus.
Fictotama maculosa^[6]	Sp. nov	Valid	Wunderlich	Miocene	Dominican amber	Dominican Republic	A tree trunk spider.
Glaesacarus^[7]	Gen. et comb. nov	Valid	Sidorchuk & Klimov	Eocene	Baltic amber	Europe (Baltic Sea region)	A mite; a new genus for "Acarus" rhombeus Koch & Berendt (1854)
Jerseyuloborus^[4]	Gen. et sp. nov	Valid	Wunderlich	Late Cretaceous	New Jersey amber	United States	A member of the family Uloboridae. Genus includes new species J. longisoma.
Macrogyion^[3]	Gen. et sp. nov		Garwood et al.	Carboniferous (late Stephanian)		France	A member of Opiliones. The type species is M. cronus.
Nephila jurassica^[8]^[9]	sp nov	Objective synonym	Selden, Shih & Ren	Middle Jurassic	Jiulongshan Formation	China	largest fossil spider described. objective syn of Mongolarachne jurassica^[9]	Mongolarachne jurassic
Labidostomma paleoluteum^[10]	Sp. nov	Valid	Dunlop & Bertrand	Eocene	Baltic amber	Europe (Baltic Sea region)	A mite, a species of Labidostomma.
Praeterleptoneta tibialis^[4]	Sp. nov	Valid	Wunderlich	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Araneomorphae belonging to the family Praeterleptonetidae. Originally described as a species of Praeterleptoneta, subsequently transferred to the separate genus Parvispina.^[11]
Scopoides dominicanus^[6]	Sp. nov	Valid	Wunderlich	Miocene	Dominican amber	Dominican Republic	A ground spider, a species of Scopoides.
Siro balticus^[12]	Sp. nov	Valid	Dunlop & Mitov	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of Opiliones, a species of Siro.
Spinocharinus^[13]	Gen. et sp. nov	Valid	Poschmann & Dunlop	Early Devonian		Germany	A member of Trigonotarbida. The type species is S. steinmeyeri.

Crustaceans

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Dynamenella miettoi^[14]	Sp. nov	Valid	De Angeli & Lovato	Eocene (Lutetian)		Italy	An isopod belonging to the family Sphaeromatidae.

Insects

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Agulla protomaculata^[15]	sp nov	valid	Engel	Lutetian	Green River Formation	USA	first snakefly from the Green River Formation
Alicodoxa^[16]	Gen et sp nov	Valid	Emeljanov & Shcherbakov	late Eocene	Rovno amber	Ukraine	A planthopper, type species A. rasnitsyni also found in Baltic amber	Alicodoxa rasnitsyni
Anonychomyrma geinitzi^[17]	Comb nov	valid	(Mayr)	Lutetian	Baltic amber	Europe	Fossil Dolichoderin ant	Anonychomyrma geinitzi
Aradus andancensis^[18]	Sp nov	valid	Marchal, Guilbert, Brisac, & Nel	Turolian	Montagne d'Andance	France	A flat bug	Aradus andancensis
Arostropsis^[19]	Gen et sp nov	valid	Yunakov & Kirejtshuk	Eocene	Baltic amber	Europe	Monotypic broad-nosed weevil genus, one species A. groehni	Arostropsis groehni
Astreptolabis^[20]	Subfam, Gen et sp nov	valid	Engel	Albian	Burmese amber	Myanmar	Monotypic earwig subfamily, one species A. ethirosomatia	Astreptolabis ethirosomatia
Baltimartyria rasnitsyni^[21]	sp nov	valid	Mey	Lutetian	Baltic amber	Europe	the second species in Baltimartyria	Baltimartyria rasnitsyni
Deinodryinus velteni^[22]	sp nov	Valid	Guglielmino & Olmi	Lutetian	Baltic amber	Europe	A dryinid wasp.	Deinodryinus velteni
Diochus electrus^[23]	sp nov	valid	Chatzimanolis & Engel	Lutetian	Baltic amber	Europe	first valid Staphylininae species from the fossil record	Diochus electrus
Dryinus rasnitsyni^[24]	sp nov	valid	Olmi & Guglielmino	Burdigalian	Dominican amber	Dominican Republic	second fossil Dryinus lamellatus group species	Dryinus rasnitsyni
Iberoraphidia^[25]	sp nov	valid	Jepsen, Ansorge, & Jarzembowski	Lower Barremian	Serra del Montsec	Spain	A mesoraphidiid snakefly, one species Iberoraphidia dividua
Lebanoraphidia^[26]	sp nov	valid	Bechly & Wolf-Schwenninger	Upper Neocomian	Jezzine	Lebanon	A mesoraphidiid snakefly, one species Lebanoraphidia nana
Leptolingia calonervis^[27]	sp.	Valid	Shi, Liu, & Ren	Middle Jurassic	Jiulongshan Formation	China
Leptolingia imminuta^[28]	sp.	Valid	Liu, Shi, & Ren	Middle Jurassic	Daohugou Beds	China	the smallest species known in the family Grammolingiidae
Litholingia ptesa^[27]	sp.	Valid	Shi, Yang, & Ren	Middle Jurassic	Jiulongshan Formation	China
Mickoleitia longimanus^[29]	ord., fam., gen. et sp.	valid	Staniczek, Bechly, & Godunko	Lower Cretaceous	Crato Formation	Brazil	A new genus in the new family Mickoleitiidae and a new insect order Coxoplectoptera.	Mickoleitia longimanus
Myanmymar^[30]	gen et sp nov	valid	Huber	Albian	Burmese amber	Myanmar	oldest mymarid described	Myanmymar aresconoides
Nanotermes^[31]	Gen et sp. nov	valid	Engel & Grimaldi	Ypresian	Cambay amber	India	A Termitid termite	Nanotermes isaacae
Ordralfabetix^[32]	gen et sp.	valid	Szwedo	Ypresian	Oise amber	France	Second fossil lophopid planthopper from Europe	Ordralfabetix sirophatanis
Parastylotermes krishnai^[31]	Sp. nov	valid	Engel & Grimaldi	Ypresian	Cambay amber	India	A Stylotermitid termite	Parastylotermes krishnai
Platyperla marquati^[33]	Species	Valid	Gallego et al.	Late Triassic	Potrerillos Formation	Argentina	A platyperlid stonefly, a species of Platyperla.
Plumalexius^[34]	fam, gen et sp.	valid	Brothers	Turonian	New Jersey amber	USA	sister genus to Plumariidae	Plumalexius rasnitsyni
Pristomyrmex rasnitsyni^[35]	sp nov	valid	Dlussky & Radchenko	Late Eocene	Scandinavian amber	Denmark	A fossil ant
Prostylotermes^[31]	Gen et sp. nov	valid	Engel & Grimaldi	Ypresian	Cambay amber	India	A Stylotermitid termite	Prostylotermes kamboja
Renphasma^[36]	Gen. et sp.	Valid	Nel & Delfosse	Early Cretaceous	Yixian Formation	China	A stick insect	Renphasma sinica
Termitaradus dominicanus^[30]	Sp nov	valid	Poinar	Burdigalian?	Dominican amber	Dominican Republic	A termitaphidid termite bug
Titanomyrma lubei^[37]	Gen. et sp.	valid	Archibald, Johnson, Mathewes, & Greenwood	Early Eocene	Green River Formation USA Messel shales, Germany Eckfeld Maar, Germany	USA Germany	new genus for the largest known ants	Titanomyrma lubei
Tytthodiplatys^[20]	Gen et sp. nov	Valid	Engel	Albian	Burmese amber	Myanmar	Oldest member of the family Diplatyidae	Tytthodiplatys mecynocercus
Undulopsychopsis^[38]	Gen et sp. nov	Valid	Peng, Makarkin, Wang, & Ren	Lower Barremian to Upper Aptian	Yixian Formation	China	A psychopsid silky lacewing, only species U. alexi	Undulopsychopsis alexi
Zophotermes^[31]	Gen et sp. nov	valid	Engel & Singh	Ypresian	Cambay amber	India	A Rhinotermitid termite	Zophotermes ashoki

Trilobites

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Chasbellus^[39]	Gen. et 2 sp. nov	Valid	Loch & Taylor	Early Ordovician	Bliss Formation House Limestone Tanyard Formation Wilberns Formation	United States	A member of Asaphida belonging to the family Symphysurinidae. The type species is C. milleri; genus also includes C. repetskii.
Gladiatoria crowei^[40]	Sp. nov	Valid	Adrain, McAdams & Westrop	Ordovician (Floian)	Fillmore Limestone Garden City Formation	United States	A member of the family Bathyuridae.
Gladiatoria harrisi^[40]	Sp. nov	Valid	Adrain, McAdams & Westrop	Ordovician (Floian)	Fillmore Limestone	United States	A member of the family Bathyuridae.
Gladiatoria nielsenae^[40]	Sp. nov	Valid	Adrain, McAdams & Westrop	Ordovician (Floian)	Fillmore Limestone Garden City Formation	United States	A member of the family Bathyuridae.
Gladiatoria phoenixi^[40]	Sp. nov	Valid	Adrain, McAdams & Westrop	Ordovician (Floian)	Fillmore Limestone Garden City Formation	United States	A member of the family Bathyuridae.
Gladiatoria reedi^[40]	Sp. nov	Valid	Adrain, McAdams & Westrop	Ordovician (Floian)	Fillmore Limestone Garden City Formation	United States	A member of the family Bathyuridae.
Hebediscus williamsi^[41]	Sp. nov	Valid	Westrop & Landing	Early Cambrian	Brigus Formation	Canada	A member of the family Hebediscidae.
Hintzeia parafirmimarginis^[42]	Sp. nov	Valid	McAdams & Adrain	Ordovician (Floian)	Fillmore Limestone Garden City Formation	United States	A member of the family Pliomeridae.
Luhops coquus^[43]	Sp. nov	Valid	Rushton	Cambrian (Drumian)	Abbey Shale	United Kingdom
Panisaspis^[44]	Gen. et 6 sp. et comb. nov	Valid	McAdams & Adrain	Early Ordovician	Fillmore Limestone Garden City Formation Yellow Hill Limestone	United States	A member of the family Pliomeridae. The type species is P. millardensis; genus also includes new species P. deltaensis, P. loganensis, P. rancherensis, P. sevierensis and P. topscityensis, as well as "Protopliomerops" quattuor Hintze (1953).
Protopliomerella bowlesi^[45]	Sp. nov	Valid	McAdams & Adrain	Ordovician (Floian)	Fillmore Limestone	United States	A member of the family Pliomeridae.
Protopliomerella kerouaci^[45]	Sp. nov	Valid	McAdams & Adrain	Ordovician (Floian)	Fillmore Limestone Garden City Formation Yellow Hill Limestone	United States	A member of the family Pliomeridae.
Protopliomerella okeeffeae^[45]	Sp. nov	Valid	McAdams & Adrain	Ordovician (Floian)	Fillmore Limestone Garden City Formation Yellow Hill Limestone	United States	A member of the family Pliomeridae.
Protopliomerella seegeri^[45]	Sp. nov	Valid	McAdams & Adrain	Ordovician (Floian)	Fillmore Limestone Garden City Formation	United States	A member of the family Pliomeridae.
Protopliomerella stegneri^[45]	Sp. nov	Valid	McAdams & Adrain	Ordovician (Floian)	Fillmore Limestone	United States	A member of the family Pliomeridae.
Psalikilopsis newmani^[46]	Sp. nov	Valid	Adrain et al.	Ordovician (Floian)	Fillmore Limestone	United States	A member of the family Bathyuridae.
Psalikilopsis paracuspidicauda^[46]	Sp. nov	Valid	Adrain et al.	Ordovician (Floian)	Fillmore Limestone Yellow Hill Limestone	United States	A member of the family Bathyuridae.
Psalikilopsis redfordi^[46]	Sp. nov	Valid	Adrain et al.	Ordovician (Floian)	Fillmore Limestone Garden City Formation	United States	A member of the family Bathyuridae.
Symphysurina ethingtoni^[39]	Sp. nov	Valid	Loch & Taylor	Late Cambrian and Early Ordovician	House Limestone	United States	A member of Asaphida belonging to the family Symphysurinidae.
Symphysurina straatmannae^[39]	Sp. nov	Valid	Loch & Taylor	Late Cambrian and Early Ordovician	Bliss Formation House Limestone Wilberns Formation	United States	A member of Asaphida belonging to the family Symphysurinidae.

References

↑ Prabhat Kumar; Neerja Jha; D.D. Bhattacharya; A.C. Pande (2011). "Acarid mites from Early Permian sediments of the Chamba valley, Himachal Pradesh, India" (PDF). Journal of the Palaeontological Society of India. 56 (2): 171–179.
↑ Ekaterina A. Sidorchuk (2018). "Mites as fossils: forever small?". International Journal of Acarology. in press. doi:10.1080/01647954.2018.1497085.
1 2 Russell J. Garwood; Jason A. Dunlop; Gonzalo Giribet; Mark D. Sutton (2011). "Anatomically modern Carboniferous harvestmen demonstrate early cladogenesis and stasis in Opiliones". Nature Communications. 2: Article number 444. doi:10.1038/ncomms1458. PMID 21863011.
1 2 3 4 Jörg Wunderlich (2011). "Some fossil spiders (Araneae) in Cretaceous ambers". In Jörg Wunderlich. Beiträge zur Araneologie, 6. pp. 539–557.
↑ Wilson R. Lourenço; Alex Beigel (2011). "A new scorpion fossil from the Cretaceous amber of Myanmar (Burma). New phylogenetic implications". Comptes Rendus Palevol. 10 (8): 635–639. doi:10.1016/j.crpv.2011.08.001.
1 2 3 4 Jörg Wunderlich (2011). "Some fossil spiders in Dominican amber (Araneae: Hersiliidae, Theridiidae, Gnaphosidae)". In Jörg Wunderlich. Beiträge zur Araneologie, 6. pp. 461–471.
↑ Ekaterina A. Sidorchuk; Pavel B. Klimov (2011). "Redescription of the mite Glaesacarus rhombeus (Koch & Berendt, 1854) from Baltic amber (Upper Eocene): evidence for female-controlled mating". Journal of Systematic Palaeontology. 9 (2): 183–196. doi:10.1080/14772019.2011.566585.
↑ Selden, P. A.; Shih, C.K.; Ren, D. (2011). "A golden orb-weaver spider (Araneae: Nephilidae: Nephila) from the Middle Jurassic of China". Biology Letters. 7 (5): 775–8. doi:10.1098/rsbl.2011.0228. PMC 3169061. PMID 21508021.
1 2 Selden, P. A.; Shih, C.K.; Ren, D. (2013). "A giant spider from the Jurassic of China reveals greater diversity of the orbicularian stem group". Naturwissenschaften. 100: 1171–1181. Bibcode:2013NW....100.1171S. doi:10.1007/s00114-013-1121-7. PMC 3889289.
↑ Jason A. Dunlop; Michel Bertrand (2011). "Fossil labidostomatid mites (Prostigmata: Labidostommatidae) from Baltic amber". Acarologia. 51 (2): 191–198. doi:10.1051/acarologia/20112006.
↑ Jörg Wunderlich (2015). "On the evolution and the classification of spiders, the Mesozoic spider faunas, and descriptions of new Cretaceous taxa mainly in amber from Myanmar (Burma) (Arachnida: Araneae)". In Jörg Wunderlich. Beiträge zur Araneologie, 9: Mesozoic spiders and other fossil arachnids. pp. 21–408.
↑ Jason A. Dunlop; Plamen G. Mitov (2011). "The first fossil cyphophthalmid harvestman from Baltic amber" (PDF). Arachnologische Mitteilungen. 40: 47–54.
↑ Markus Poschmann; Jason A. Dunlop (2011). "Trigonotarbid arachnids from the Lower Devonian (Siegenian) of Bürdenbach (Lahrbach Valley, Westerwald area, Rhenish Slate Mountains, Germany)". Paläontologische Zeitschrift. 85 (4): 433–447. doi:10.1007/s12542-011-0109-x.
↑ Antonio De Angeli; Alessandro Lovato (2011). "Dynamenella miettoi n. sp., nuovo isopode (Crustacea, Flabellifera, Sphaeromatidae) dell'Eocene del Veneto (Vicenza, Italia settentrionale)" (PDF). Lavori – Società Veneziana di Scienze Naturali. 36: 111–118.
↑ Engel, Michael S. (2011). "A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae)". Transactions of the Kansas Academy of Science. 114 (1–2): 77–87. doi:10.1660/062.114.0107.
↑ Emeljanov, A. F.; Shcherbakov, D. E. (2011). "A new genus and species of Dictyopharidae (Homoptera) from Rovno and Baltic amber based on nymphs". ZooKeys. 130: 175–184. doi:10.3897/zookeys.130.1775.
↑ Heterick, B. E.; Shattuck, S. (2011). "Revision of the ant genus Iridomyrmex (Hymenoptera: Formicidae)". Zootaxa. 2845: 169.
↑ Marchal, L.; Guilbert, E.; Brisac, P.; Nel., A. (2011). "A new record and a new species of Aradidae fossils (Hemiptera: Heteroptera)". Zootaxa. 2832: 56–62.
↑ Yunakov, N.N.; Kirejtshuk, A.G. (2011). "New genus and species of broad-nosed weevils from Baltic amber and notes on fossils of the subfamily Entiminae (Coleoptera, Curculionidae)". ZooKeys. 160: 73–96. doi:10.3897/zookeys.160.2108. PMC 3253632. PMID 22303121.
1 2 Engel, M.S. (2011). "New earwigs in mid-Cretaceous amber from Myanmar (Dermaptera, Neodermaptera)". ZooKeys. 130: 137–152. doi:10.3897/zookeys.130.1293. PMC 3260755. PMID 22259272.
↑ Mey, Wolfram (2011). "On the systematic position of Baltimartyria Skalski, 1995 and description of a new species from Baltic amber (Lepidoptera, Micropterigidae)". ZooKeys. 130: 331–342. doi:10.3897/zookeys.130.1480. PMC 3260770. PMID 22259287.
↑ Guglielmino, A.; Olmi, M. (2011). "Revision of fossil species of Deinodryinus, with description of a new species (Hymenoptera, Dryinidae)". ZooKeys. 130: 495–504. doi:10.3897/zookeys.130.1326. PMC 3260777. PMID 22259295.
↑ Chatzimanolis, S.; Engel, M. S. (2011). "A new species of Diochus from Baltic amber (Coleoptera, Staphylinidae, Diochini)" (PDF). ZooKeys. 138: 65–73. doi:10.3897/zookeys.138.1896. PMC 3208521. PMID 22144854.
↑ Olmi, Massimo; Guglielmino, Adalgisa (2011). "Revision of fossil species of Dryinus belonging to lamellatus group, with description of a new species (Hymenoptera, Dryinidae)" (PDF). 130: 505–514. doi:10.3897/zookeys.130.133.
↑ Jepson, J.E.; Ansorge, J.; Jarzembowski, E.A. (2011). "New snakeflies (Insecta: Raphidioptera) from the Lower Cretaceous of the UK, Spain and Brazil". Palaeontology. 54 (2): 385–395. doi:10.1111/j.1475-4983.2011.01038.x.
↑ Bechly, G.; Wolf-Schwenninger, K. (2011). "A new fossil genus and species of snakefly (Raphidioptera: Mesoraphidiidae) from Lower Cretaceous Lebanese amber, with a discussion of snakefly phylogeny and fossil history" (PDF). Insect Systematics and Evolution. 42 (2): 221–236. doi:10.1163/187631211X568164.
1 2 Shi, C.; Yang, Q.; Ren, D. (2011). "Two new fossil lacewing species from the Middle Jurassic of Inner Mongolia, China (Neuroptera: Grammolingiidae)" (PDF). Acta Geologica Sinica. 85: 482–489. doi:10.1111/j.1755-6724.2011.00416.x. Archived from the original (PDF) on 2012-03-24.
↑ Liu, Y.; Shi, C.; Ren, D. (2011). "A new lacewing (Insecta: Neuroptera: Grammolingiidae) from the Middle Jurassic of Inner Mongolia, China" (PDF). Zootaxa. 2897: 51–56.
↑ Staniczek, A.H.; Bechly, G.; Godunko, R.J. (2011). "Coxoplectoptera, a new fossil order of Palaeoptera (Arthropoda: Insecta), with comments on the phylogeny of the stem group of mayflies (Ephemeroptera)" (PDF). Insect Systematics & Evolution. 42 (2): 101–138. doi:10.1163/187631211X578406.
1 2 Poinar, G.; Huber, J. T. (2011). "A new genus of fossil Mymaridae (Hymenoptera) from Cretaceous amber and key to Cretaceous mymarid genera". ZooKeys. 130: 461–472. doi:10.3897/zookeys.130.1241. PMC 3260775. PMID 22259293.
1 2 3 4 Engel, M.S.; Grimaldi, D.A.; Nascimbene, P.C.; Singh, H. (2011). "The termites of Early Eocene Cambay amber, with the earliest record of the Termitidae (Isoptera)". ZooKeys. 148: 105–123. doi:10.3897/zookeys.148.1797. PMC 3264413. PMID 22287892.
↑ Szwedo, J. (2011). "Ordralfabetix sirophatanis gen. et sp. n.—the first Lophopidae from the Lowermost Eocene Oise amber, Paris Basin, France (Hemiptera: Fulgoromorpha)" (PDF). Zootaxa. 2822: 52–60.
↑ Gallego, O.F.; Rébori, L; Zavattieri, A.M.; Sinitshenkova, N.; Lara, M.B. ,; Martins-Neto, R.G. (2011). "The most ancient Platyperlidae (Insecta, Perlida= Plecoptera) from early Late Triassic deposits in southern South America". Ameghiniana. 48 (4): 447–461. doi:10.5710/amgh.v48i4(360).
↑ Brothers, Denis J. (2011). "A new Late Cretaceous family of Hymenoptera, and phylogeny of the Plumariidae and Chrysidoidea (Aculeata)" (PDF). ZooKeys. 130: 515–542. doi:10.3897/zookeys.130.1591. PMC 3260779. PMID 22259297.
↑ Dlussky, G. M.; Radchenko, A. G. (2011). "Pristomyrmex rasnitsyni sp. n., the first known fossil species of the ant genus Pristomyrmex Mayr (Hymenoptera: Formicidae) from the Late Eocene Danish amber". Russian Entomological Journal. 20 (3): 251–254.
↑ Nel, A.; Delfosse, E. (2011). "A new Chinese Mesozoic stick insect". Acta Palaeontologica Polonica. 56 (2): 429–432. doi:10.4202/app.2009.1108.
↑ Archibald, S. Bruce; Johnson, Kirk R.; Mathewes, Rolf W.; Greenwood, David R. (2011). "Intercontinental dispersal of giant thermophilic ants across the Arctic during early Eocene hyperthermals". Proceedings of the Royal Society B. 278 (1725): 3679–3686. doi:10.1098/rspb.2011.0729. PMC 3203508. PMID 21543354.
↑ Peng, Y.; Makarkin, V.N.; Wang, X.; Ren, D. (2011). "A new fossil silky lacewing genus (Neuroptera, Psychopsidae) from the Early Cretaceous Yixian Formation of China". ZooKeys. 130: 217–228. doi:10.3897/zookeys.130.1576. PMC 3260761. PMID 22259278.
1 2 3 James D. Loch; John F. Taylor (2011). "New symphysurinid trilobites from the Cambrian-Ordovician boundary interval in the western United States". Memoirs of the Association of Australasian Palaeontologists. 42: 417–436.
1 2 3 4 5 Jonathan M. Adrain; Neo E.B. McAdams; Stephen R. Westrop (2011). "Affinities of the Lower Ordovician (Tulean; lower Floian) trilobite Gladiatoria, with species from the Great Basin, western United States" (PDF). Memoirs of the Association of Australasian Palaeontologists. 42: 321–367.
↑ Stephen R. Westrop; Ed Landing (2011). "Lower Cambrian (Branchian) eodiscoid trilobites from the lower Brigus formation, Avalon Peninsula, Newfoundland, Canada". Memoirs of the Association of Australasian Palaeontologists. 42: 209–262.
↑ Neo E.B. McAdams; Jonathan M. Adrain (2011). "Systematics of the Lower Ordovician pliomerid trilobite Hintzeia, with species from the Great Basin, western USA" (PDF). Zootaxa. 2910: 1–45.
↑ Adrian W.A. Rushton (2011). "The mid-Cambrian (Drumian) centropleurid trilobite Luhops and its relatives from the Abbey Shale formation near Nuneaton, central England". Memoirs of the Association of Australasian Palaeontologists. 42: 93–104.
↑ Neo E.B. McAdams; Jonathan M. Adrain (2011). "Panisaspis, a new genus of pliomerid trilobites from the Lower Ordovician (Ibexian; Tulean and Blackhillsian) of the Great Basin, western USA" (PDF). Zootaxa. 2969: 1–68.
1 2 3 4 5 Neo E.B. McAdams; Jonathan M. Adrain (2011). "Revision of the Lower Ordovician (lower Floian; Tulean) pliomerid trilobite Protopliomerella, with new species from the Great Basin, western USA" (PDF). Zootaxa. 3144: 1–113.
1 2 3 Jonathan M. Adrain; Neo E.B. McAdams; Stephen R. Westrop; Talia S. Karim (2011). "Systematics and affinity of the Lower Ordovician (Tulean; lower Floian) trilobite Psalikilopsis" (PDF). Memoirs of the Association of Australasian Palaeontologists. 42: 369–416.

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[44] Neo E.B. McAdams; Jonathan M. Adrain (2011). "Panisaspis, a new genus of pliomerid trilobites from the Lower Ordovician (Ibexian; Tulean and Blackhillsian) of the Great Basin, western USA" (PDF). Zootaxa. 2969: 1–68.

[ZTProtopliomerella-45] 1 2 3 4 5 Neo E.B. McAdams; Jonathan M. Adrain (2011). "Revision of the Lower Ordovician (lower Floian; Tulean) pliomerid trilobite Protopliomerella, with new species from the Great Basin, western USA" (PDF). Zootaxa. 3144: 1–113.

[AdrainetalPsalikilopsis-46] 1 2 3 Jonathan M. Adrain; Neo E.B. McAdams; Stephen R. Westrop; Talia S. Karim (2011). "Systematics and affinity of the Lower Ordovician (Tulean; lower Floian) trilobite Psalikilopsis" (PDF). Memoirs of the Association of Australasian Palaeontologists. 42: 369–416.