← Lycophytes |
Flora of New York — Pre-polypod ferns | → Polypodiales | |||
Table of contents |
Genus index |
Protected species index | Invasive species index | ||
Order Ophioglossales
The order Ophioglossales contains the oldest of the plants now considered to be ferns s.l.[1]
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Family Ophioglossaceae
The Ophioglossaceae (adder's-tongue family) comprises primitive ferns that have traditionally been considered part of a group informally known as fern-allies. In New York, the family is represented by about a dozen native species, all of which are listed as protected plants.[1]
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Botrychium
The genus Botrychium contains the moonworts, evergreen grapeferns, and rattlesnake ferns, each of which is treated here as a subgenus of Botrychium. These groups are also sometimes treated as separate genera within the Ophioglossaceae.
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Botrychium subg. Botrychium
Botrychium subg. Botrychium | Moonwort, grape fern | N.Y. Status | Images | Distribution | NY NPT |
(Döll) A.Braun ex W.D.J.Koch 1843. B. lunaria var. matricariifolium 1847. B. neglectum Alph.Wood 1847. B. matricariifolium ex W.D.J.Koch |
Daisy-leaved moonwort, Chamomile grape-fern, Matricary grapefern Botryche à feuille de matricaire |
Native, C:7, Likely secure FACU Perennial, Fern |
NYFA-4 USDA-NN NPT Can FNA BNA Images, wsp | ||
(Pease & A.H.Moore) Fernald 1906. B. lanceolatum var. angustisegmentum 1915. B. angustisegmentum (Pease & A.H.Moore) Fernald 1937. B. lanceolatum ssp. angustisegmentum |
Narrow triangle moonwort, Narrow moonwort, Lance-leaved grape-fern Botryche à segments étroits |
Native, C:7, Likely secure FACW Perennial, Fern |
NYFA-4 USDA-NN NPT Can ITIS FNA Trop. BNA Images, wsp | ||
E.Hitchc. 1823. B. simplex E.Hitchc. 1899. B. tenebrosum A.A.Eaton |
Least grape-fern, Little grape-fern |
Native, C:9, Likely secure Perennial, Fern |
NYFA-4-5 USDA-NN NPT BNA Images, wsp | ||
A.A.Eaton B. simplex var. tenebrosum B. tenebrosum A.A.Eaton |
Shade-loving moonwort, Swamp moonwort |
Native, C:10, Endangered Perennial, Fern |
NYFA-U NPT Images, wsp | ||
Stensvold & Farrar 1753. Osmunda lunaria L. misapplied
1801. B. lunaria (L.) Sw. misappl.
1903. B. onondagense Underw.
1923. B. lunaria var. onondagense
2017. B. neolunaria Stensvold & Farrar
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Moonwort grape-fern, Common moonwort |
Native, C:10, Endangered, CA, NYNHP: 1[1] Perennial, Fern |
NYFA-1 USDA-NN NPT Can FNA BNA Images, wsp | ||
Vict. 1927. B. minganense Vict. 1937. B. lunaria var. minganense 1965. B. lunaria ssp. minganense |
Mingan's moonwort, Mingan moonwort, Mingan Island moonwort Botryche de Mingan |
Native, C:10, Endangered, CA, NYNHP: 1[2] Perennial, Fern |
NYFA-1 USDA-N NPT BNA Images, wsp | ||
W.H.Wagner & Farrar B. minganense (in part) |
Prairie moonwort, Prairie dunewort, Dunewort, Iowa moonwort |
Native, C:10, No recent reports, NYNHP: 1[3] Perennial, Fern |
NYFA-Z USDA-NN NPT BNA Images, wsp | ||
Botrychium subg. Botrychium (excluded taxa) | Moonwort, grape fern | N.Y. Status | Images | Distribution | NY NPT |
(S.G.Gmel.) Ångstr. 1854. B. lanceolatum (S.G.Gmel.) Ångstr. |
Triangle moonwort Botryche élancé |
N. America native, N.Y. excluded Perennial, Fern |
NYFA-XCLD FNA Images, wsp | ||
W.H.Wagner 1986. B. ascendens W.H.Wagner |
Triangle-lobe moonwort, Upswept moonwort, Ascending grapefern Botryche ascendant |
Native, Unreported Perennial, Fern |
NYFA-D USDA-NN NPT Can ITIS FNA Trop. BNA Images, wsp | ||
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Botrychium subg. Osmundopteris
The subgenus Osmundopteris (sometimes treated as the genus Botrypus) contains a single, fairly widespread Rattlesnake fern species in New York.
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Botrychium subg. Osmundopteris | Rattlesnake fern | N.Y. Status | Images | Distribution | NY NPT |
(L.) Sw. 1753. Osmunda virginiana L.
1800. Botrychium virginianum Sw.
1803. Botrypus virginianus Michx.
1814. Botrychium gracile Pursh
1938. Osmundopteris virginiana Small
1958. Japanobotrychium virginianum M.Nishida
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Rattlesnake fern, Rattlesnake grapefern, Virginia grapefern Botryche de Virginie |
Native, C:8, Secure |
NYFA-5 USDA-NN NPT Can FNA Trop. Images, wsp | ||
Botrychium subg. Sceptridium
Botrychium subg. Sceptridium | Evergreen grape fern | N.Y. Status | Images | Distribution | NY NPT |
Spreng. 1804. Botrychium dissectum Spreng. 1810. Botrychium obliquum Muhl. 1902. B. dissectum var. obliquum 1905. Sceptridium dissectum Lyon 1905. Sceptridium obliquum Lyon |
Cut-leaved grape-fern, Cutleaf grapefern, Dissected grapefern |
Native, C:3, Secure |
NYFA-5 USDA-NN NPT Can FNA Trop. BNA Images, wsp Note: CA | ||
(S.G.Gmel.) Rupr. B. californicum Underw. B. coulteri Underw. B. silaifolium C.Presl Sceptridium multifidum |
Leathery grape-fern, Leather grape-fern |
Native, C:3, Secure |
NYFA-5 USDA-NN NPT BNA Images, wsp | ||
(Gilbert) House 1901. B. ternatum var. oneidense 1905. B. oneidense House 1916. B. multifidum var. oneidense 1924. B. dissectum var. oneidense 1998. Sceptridium oneidense Holub |
Blunt-lobed grape-fern, Bluntlobe grapefern, Oneida grape fern |
Native, C:5, Threatened, CA, NYNHP: 2[1] |
NYFA-2-3 USDA-NN NPT BNA Images, wsp | ||
W.H.Wagner B. multifidum fo. dentatum auct. B. ternatum non (Thunb.) Sw. Sceptridium rugulosum |
Ternate grapefern, Rugulose grape-fern, St. Lawrence grape-fern, Botryche à limbe rugueux |
Native, C:6, Endangered, Few populations, CA, NYNHP: 1[2] |
NYFA-1 USDA-NN NPT FNA BNA Images, wsp | ||
Botrychium subg. Sceptridium (excluded taxa) | Evergreen grape fern | N.Y. Status | Images | Distribution | NY NPT |
(Savigny) Underw. 1798. Osmunda biternata Savigny 1896. B. biternatum (Savigny) Underw. 1903. B. tenuifolium Underw. 1905. Sceptridium tenuifolium (Underw.) Lyon 1905. S. biternatum (Savigny) Lyon 1924. B. dissectum var. tenuifolium |
Sparselobe grapefern, Sparse-lobe grape fern |
N. America native, N.Y. excluded |
NYFA-XCLD USDA-N0 FNA Trop. Images, wsp | ||
Ophioglossum
Ophioglossum | Adder's-tongue | N.Y. Status | Images | Distribution | NY NPT |
Raf. 1814. Ophioglossum pusillum Raf. 1927. O. vulgatum var. alaskanum 1917. O. vulgatum var. pseudopodum auct. O. vulgatum non L. (s.s.) |
Northern adder's-tongue Herbe-sans-couture |
Native, C:5, Vulnerable FACW Perennial, Fern |
NYFA-3 USDA-N NPT ITIS FNA IPN LBJ Images, wsp | ||
Ophioglossum (excluded) | Adder's-tongue | N.Y. Status | Images | Distribution | NY NPT |
L. 1753. Ophioglossum vulgatum L. 1939. O. vulgatum var. pycnostichum 1977. O. pycnostichum (Fernald) A.Löve & D.Löve |
Southern adder's-tongue, English adder's-tongue Herbe sans couture |
N. America native, N.Y. excluded |
NYFA-XCLD USDA-N0 ARS FNA IPN LBJ Images, wsp | ||
Order Equisetales
The order Equisetales contains the plants commonly known as horsetails and scouring rushes. This order contains the single living genus Equisetum, which is represented in every county of New York.
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Family Equisetaceae
The Equisetaceae (horsetail family) ...
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Equisetum
Equisetum is here divided into the two subgenera Equisetum (horsetails) and Hippochaete (scouringrushes).
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Equisetum subg. Equisetum
Equisetum subg. Equisetum | Horsetail | N.Y. Status | Images | Distribution | NY NPT |
L. 1753. Equisetum arvense L. 1832. Equisetum boreale Bong. 1912. Allostelites arvensis (L.) Börner 1960. Equisetum calderi B.Boivin |
Field horsetail, Common horsetail, Common field horsetail Prêle des champs, Queue de renard |
Native, C:2, Secure FAC |
NYFA-5 USDA-NN NPT Can ARS ITIS FNA Trop. NSE IPN Images, wsp | ||
L. 1753. Equisetum fluviatile L. 1753. Equisetum limosum L. 1904. E. fluviatile var. limosum (L.) Farw. |
Water horsetail, River horsetail Prêle fluviatile |
Native, C:4, Secure OBL |
NYFA-5 USDA-NN NPT ARS ITIS FNA Trop. Images, wsp | ||
Kühlew. ex Rupr. (pro sp.) Equisetum arvense × Equisetum fluviatile 1845. Equisetum litorale Kühlew. ex Rupr. |
Litoral horsetail Prêle littorale |
Native, Rare OBL |
NYFA-3 USDA-NN NPT ITIS FNA Images, wsp | ||
L. 1753. Equisetum sylvaticum L. 1795. Equisetum capillare Hoffm. |
Wood horsetail, Woodland horsetail Prêle des bois |
Native, C:7, Secure FACW |
NYFA-5 USDA-NN NPT ARS ITIS FNA Images, wsp | ||
L. 1753. Equisetum palustre L. 1927. E. palustre var. americanum Vict. |
Marsh horsetail Prêle des marais |
Native, C:5, Threatened, NYNHP: 2[1] FACW |
NYFA-2 USDA-NN NPT ARS ITIS FNA Images, wsp | ||
Ehrh. 1784. Equisetum pratense Ehrh. |
Meadow horsetail, Shady horsetail Prêle des prés |
Native, C:4, Threatened, NYNHP: 2[2] FACW |
NYFA-2 USDA-NN NPT ARS ITIS FNA Images, wsp | ||
Equisetum (excluded) subg. Equisetum | Horsetail | N.Y. Status | Images | Distribution | NY NPT |
Ehrh. ssp. braunii (Milde) Hauke 1862. Equisetum braunii Milde 1978. Equisetum telmateia var. braunii auct. Equisetum maximum non Lam. |
Giant horsetail, Great horsetail, Ivory horsetail Prêle géante |
Introduced from western N. America, N.Y. excluded OBL |
NYFA-XCLD USDA-NN Can ARS ITIS FNA Images, wsp | ||
Equisetum subg. Hippochaete
Christenhusz et. al. (2019) suggests that the western scouringrush (Equisetum hyemale ssp. affine) should be treated at the species level (Equisetum praealtum Raf.).[1]
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Equisetum subg. Hippochaete | Scouringrush | N.Y. Status | Images | Distribution | NY NPT |
L. ssp. affine Calder & Roy L.Taylor Equisetum affine Engelm. Equisetum praealtum Raf. Equisetum robustum A.Braun Hippochaete hyemalis (L.) Braun |
Common scouringrush, Western scouringrush, Rough horsetail, Scouringrush horsetail Prêle d'hiver |
Native, C:2, Secure |
NYFA-5 USDA-NN NPT iNat ARS ITIS FNA Images, wsp | ||
Michx. (1803) |
Dwarf scouring rush Prêle faux-scirpe |
Native, C:9, Secure |
NYFA-5 USDA-NN ITIS FNA Images, wsp | ||
Schleich. ex F.Weber & D.Mohr ssp. variegatum Equisetum variegatum Schleich. ex F.Weber & D.Mohr Equisetum variegatum var. variegatum Hippochaete variegata (Schleich. ex F.Weber & D.Mohr) Bruhin |
Variegated scouring rush, Variegated horsetail Prêle panachée |
Native, C:3, Likely secure FACW Perennial, Herb |
NYFA-4 USDA-NN NPT Can ITIS FNA Images, wsp | ||
Equisetum hyemale × Equisetum variegatum |
Rough-toothed scouring rush | Native, C:3, Vulnerable |
NYFA-3 USDA-N Images, wsp | ||
(A.A.Eaton) J.H.Schaffn. Equisetum laevigatum × Equisetum variegatum 1904. E. variegatum var. nelsonii 1926. E. nelsoni J.H.Schaffn. |
Nelson's scouring rush, Nelson's horsetail |
Native, Unranked |
NYFA-U USDA-NN ARS ITIS FNA Images, wsp | ||
Clute Equisetum hyemale × Equisetum laevigatum 1903. E. hyemale var. intermedium 1904. Equisetum ferrissii Clute 1917. Equisetum intermedium Rydb. 1951. E. hyemale var. elatum |
Ferris' scouring rush | Native, Unranked |
NYFA-U USDA-NN ARS Images, wsp | ||
Equisetum (excluded) subg. Hippochaete | Scouring rush | N.Y. Status | Images | Distribution | NY NPT |
A.Braun (1844) E. funstonii E. kansanum Hippochaete laevigata |
Smooth scouring-rush | N. America native, N.Y. excluded |
NYFA-XCLD USDA-NN Images, wsp | ||
Order Osmundales
The Osmundales contains a single family with three native species represented outside of cultivation in New York.
Family Osmundaceae
The Osmundaceae (royal fern family)...
Osmundastrum
Osmundastrum | Cinnamon fern | N.Y. Status | Images | Distribution | NY NPT |
(L.) C.Presl var. cinnamomeum 1753. Osmunda cinnamomea L. 1841. Osmunda cinnamomea var. frondosa 1847. Osmundastrum cinnamomeum (L.) C.Presl 1868. Osmunda cinnamomea var. imbricata 1902. Osmunda cinnamomea var. glandulosa 1909. Osmunda cinnamomea var. fokiensis |
Cinnamon fern, Buckhorn Osmonde cannelle |
Native, C:6, Secure, DEC-D[1] OBL-FACW Perennial, Herb-fern |
NYFA-5 USDA-NN NPT Can ARS ITIS Trop. Images, wsp Note: FEIS | ||
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Osmunda
Osmunda subg. Claytosmunda | Flowering fern | N.Y. Status | Images | Distribution | NY NPT |
L. 1753. Osmunda claytoniana L. 1801. Struthiopteris claytoniana (L.) Bernh. 1848. Plenasium claytonianum (L.) C. Presl 1941. Osmundastrum claytonianum (L.) Tagawa |
Interrupted fern, Clayton's fern Osmonde de Clayton |
Native, C:4, Secure, DEC-D[1] FAC Perennial, Herb-fern |
NYFA-5 USDA-N NPT Can Trop. Images, wsp | ||
Osmunda subg. Osmunda | Flowering fern | N.Y. Status | Images | Distribution | NY NPT |
L. var. spectabilis (Willd.) A. Gray 1810. Osmunda spectabilis Willd. 1856. Osmunda regalis var. spectabilis 1977. Osmunda regalis ssp. spectabilis |
Royal fern, Regal fern, Flowering fern Osmonde royale, Fougère royale |
Native, C:6, Secure, DEC-D[1] OBL Perennial, Herb-fern |
NYFA-5 USDA-N Can ARS Images, wsp | ||
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Order Hymenophyllales
The order Hymenophyllales contains a single family, the Hymenophyllaceae, or filmy fern family. In New York, only one species of this family has been verified as living outside of cultivation, and that species is considered to be endangered in the state.
Family Hymenophyllaceae
Crepidomanes
The only member of the Hymenophyllaceae found in New York is a gametophitic weft fern Crepidomanes intricatum, which was transferred from Trichomanes to the mostly Asian Crepidomanes genus by Ebihara & Weakley (2011) based on molecular sequence. Crepidomanes intricatum is assumed to be gametophitic only. Sporophitic generations of this plant have not been observed.[1][2]
Crepidomanes | Grotto felt | N.Y. Status | Images | Distribution | NY NPT |
(Farrar) Ebihara & Weakley 1992. Trichomanes intricatum Farrar 2011. Crepidomanes intricatum (Farrar) Ebihara & Weakley |
Appalachian bristle fern, Appalachian trichomanes, Filmy fern, Weft fern |
Native, C:10, Endangered, DEC-A[3] |
NYFA-1 USDA-N0 NPT ITIS FNA Trop. Images, wsp | ||
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Order Schizaeales
The order Schizaeales contains the seemingly dissimilar curly-grass and climbing ferns.
Family Lygodiaceae
The Lygodiaceae (climbing-fern family) consists of the single genus Lygodium, which in New York, is represented by the endangered American climbing fern.
Lygodium
Lygodium | Climbing fern | N.Y. Status | Images | Distribution | NY NPT |
(Bernh.) Sw. 1800. Gisopteris palmata Bernh. 1806. Lygodium palmatum (Bernh.) Sw. |
American climbing fern, Creeping fern, Hartford fern, Windsor fern |
Native, C:10, Endangered, DEC-A[1] FACW Perennial, Herb-fern |
NYFA-1 USDA-N0 FNA Images, wsp | ||
Family Schizaeaceae
The Schizaeaceae (curly-grass family)...
Schizaea
Schizaea Sm. | Curlygrass fern | N.Y. Status | Images | Distribution | NY NPT |
Pursh 1814. Schizaea pusilla Pursh |
Curlygrass, Curly-grass fern, Little curlygrass fern |
Native, C:9, Endangered, DEC-A[1] OBL Perennial, Herb-fern |
NYFA-1 USDA-N FNA Images, wsp | ||
Order Salviniales
The order Salviniales (the heterosporous or water ferns) contains the families Salviniaceae (mosquitoferns) and Marsileaceae (waterferns), which have few species represented in New York.
Family Salviniaceae
The family Salviniaceae contains the aquatic fern genera Azolla and Salvinia.
Azolla
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Azolla sect. Azolla | Mosquitofern | N.Y. Status | Images | Distribution | NY NPT |
Kaulf. 1824. Azolla cristata Kaulf.
1827. Azolla portoricensis Spreng.
1830. Azolla mexicana Schltdl. & Cham. nom. nud.
1844. Azolla mexicana Schltdl. & Cham. ex Kunze
1845. Azolla mexicana Schltdl. & Cham. ex C.Presl
auct. Azolla caroliniana non Willd.
auct. Azolla filiculoides non Lam.
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Mosquito fern, Carolina mosquitofern, Eastern mosquito-fern, Fairy-moss, Mosquito-plant Azolle de Caroline, Azolla de Caroline |
Native, C:2, Threatened, DEC-D[1] OBL Annual, Herb-fern |
NYFA-2? USDA-NX NPT Can ITIS FNA Trop. Images, wsp Note: [2][3] | ||
Azolla (excluded taxa) | Mosquitofern | N.Y. Status | Images | Distribution | NY NPT |
Lam. 1783. Azolla filiculoides Lam.
1810. Azolla caroliniana Willd.
1810. Azolla magellanica Willd.
1824. Azolla microphylla Kaulf.
1827. Azolla arbuscula Desv.
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Pacific mosquitofern, Pacific azolla, Fern azolla, Large mosquito fern, Duckweed fern, Redwater fern Azolle fausse-filicule, Azolla fausse-filicule |
N. America native, N.Y. excluded |
NYFA-XCLD USDA-NX Can ARS ITIS FNA Trop. Images, wsp | ||
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Salvinia
The two species of Salvinia previously reported in New York have been excluded in the New York Flora Atlas as of 9/18/2016.[1]
The genus Salvinia (watermoss) contains mostly tropical, aquatic ferns. S. minima, mistakenly reported as S. natans has been reported from ponds on Staten Island and Rockland County, but these may have been aquarium escapes, as this genus is unlikely to be able to naturalize that far north.[2]
DEC regulation 193.3 Protected native plants, lists Salvinia minima as Exploitably vulnerable,[3] even though it is a Central and South American native that is considered to be invasive in the southeastern U.S.[4]
Salvinia Seg. (excluded species) | Watermoss | N.Y. Status | Images | Distribution | NY NPT |
Baker 1886. Salvinia minima Baker auct. Salvinia natans non (L.) All. auct. Salvinia rotundifolia non Willd. |
Floating-fern, Water-fern, Water spangles, Common watermoss |
Introduced from S. America, N.Y. excluded, DEC-D[5] |
NYFA-XCLD USDA-XX NPT ARS Images, wsp | ||
1753. Marsilea natans L. 1785. Salvinia natans (L.) All. |
Floating watermoss | Introduced from Eurasia, northen Africa, N.Y. excluded, Old World native |
NYFA-XCLD USDA-X ARS Images, wsp Note: [6] | ||
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Family Marsileaceae
The Marsileaceae (water-clover family) has a single species (Marsilea quadrifolia) surviving outside of cultivation in New York.
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Marsilea
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Marsilea | Waterclover | N.Y. Status | Images | Distribution | NY NPT |
European water-clover, European water fern, Water shamrock, European pepperwort Marsilée à quatre feuilles, Marsile à quatre feuilles |
Introduced from Europe, Potentially invasive, NYIS: 45%[1] |
NYFA-X USDA-XN‽ NPT Can ARS FNA Images, wsp | |||
← Lycophytes |
Flora of New York — Pre-polypod ferns | → Polypodiales | |||
Table of contents |
Genus index |
Protected species index | Invasive species index | ||