< Flora of New York

Pre-polypod
ferns
Flora of New York Polypodiales
Pinales
Table of
contents
Genus
index
Protected species index Invasive species index
The classification used here for the order Polypodiales Link (1833) follows Christenhusz & Chase (2014)[1]
  1. Maarten J.M. Christenhusz & Mark W. Chase (2014). "Trends and concepts in fern classification". Annals of Botany 113 (9): 571–594. doi:10.1093/aob/mct299.PDF
Order Family Subfamily Genus Common name(s) #
Polypodiales Dennstaedtiaceae Dennstaedtia hay-scented fern 1
Pteridium brackenfern 2
Pteridaceae Cryptogrammoideae Cryptogramma rock brake 1
Pteridoideae Pteris brake fern 1
Cheilanthoideae Pellaea cliff brake 2
Cheilanthes lip fern 1
Vittarioideae Adiantum maidenhair-fern 1
Vittaria ribbon fern 1
Aspleniaceae Cystopteridoideae Cystopteris bladder fern 5
Gymnocarpium oak fern 2
Asplenioideae Asplenium spleenwort 10
Thelypteridoideae Phegopteris beech fern 2
Thelypteris maiden fern, New York fern 3
Woodsioideae Woodsia cliff fern 5
Blechnoideae Onoclea sensitive fern 1
Matteuccia ostrich fern 1
Woodwardia chain fern 2
Athyrioideae Deparia false spleenwort 1
Homalosorus glade fern 1
Athyrium lady fern 3
Polypodiaceae Dryopteridoideae Cyrtomium Japanese hollyfern 1
Dryopteris woodfern 30
Polystichum hollyfern 5
Polypodioideae Polypodium polypody, rockcap fern 3
Pleopeltis scaly-polypody, resurrection fern 1

Family Dennstaedtiaceae

The Dennstaedtiaceae Lotsy (1909), (bracken-fern family)

Although they are native to the Northeast, the hay-scented fern (Dennstaedtia punctilobula), and brackenfern (Pteridium aquilinum) are considered to be "interfering" plants in New York forests. These ferns produce dense stands that prevent the establishment of tree seedlings. While this has been found to be beneficial in limited situations, such as under power-lines and on ski-area trails, these ferns are generally seen as detrimental to forest regeneration.[1]


Dennstaedtia

Dennstaedtia punctilobula
hay-scented fern
Hay-scented fern (Dennstaedtia punctilobula) is often considered to be an interfering species in New York forest understories, where large deer populations over-browse other native plants but avoid this fern, which prevents tree regeneration.
    Polypodiales — Dennstaedtiaceae — Dennstaedtia
    Dennstaedtia Hay-scented fern N.Y. Status Images Distribution NY NPT
    (Michx.) T.Moore

    1803. Nephrodium punctilobulumMichx.
    1810. Aspidium punctilobumWilld.
    1827. Sitobolium punctilobumDesv.
    1848. Dicksonia punctilobulaA.Gray
    1857. Dennstaedtia punctilobulaT.Moore
    Eastern hay-scented fern,
    Hay-scent fern,
    Boulder fern
    Dennstaedtie à
    lobules ponctués
    Native, C:4,
    Secure,
    Interfering

    FACU-UPL

    Perennial,
    Herb-fern
    NYFA-5
    USDA-NN
    NPT Can
    ARS ITIS
    FNA Trop.



    Images, wsp

      Pteridium

      Pteridium aquilinum
      eastern bracken fern

        Polypodiales — Dennstaedtiaceae — Pteridium
        Pteridium Brackenfern N.Y. Status Images Distribution NY NPT
        (L.) Kuhn
        var. latiusculum(Desv.) Underw. ex A. Heller

        1827. Pteris latiusculaDesv.
        1909. Pteridium aquilinum var. latiusculum
        1914. Pteridium latiusculum(Desv.) Hieron.
        1941. Pteridium aquilinum ssp. latiusculum
        Eastern brackenfern,
        Eastern bracken,
        Brackenfern
        Fougère-aigle de l'Est
        Native, C:3,
        Secure,
        Interfering

        FACU

        Perennial,
        Fern
        NYFA-5
        USDA-NN
        NPT Can
        ARS ITIS
        FNA Trop.
        BNA


        Images, wsp
        (L.) Kuhn
        var. pseudocaudatum(Clute) A. Heller

        1900. Pteris aquilina var. pseudocaudataClute
        1900. Pterid. aquilinum var. pseudocaudatum
        1919. Pterid. latiusculum var. pseudocaudatum
        1941. Pterid. aquilinum ssp. pseudocaudatum
        Southeastern Bracken,
        Tailed bracken,
        Brackenfern,
        Western brackenfern
        Native, C:3,
        Endangered,
        DEC-A[1]

        FACU

        Perennial,
        Fern
        NYFA-Z
        USDA-N0
        NPT 
        ITIS
        FNA Trop.
        BNA


        Images, wsp
        Pteridium(excluded) Brackenfern N.Y. Status Images Distribution NY NPT
        (L.) Kuhn
        ssp. aquilinum

        1753. Pteris aquilinaL.
        1799. Asplenium aquilinum(L.) Bernh.
        1879. Pteridium aquilinum(L.) Kuhn
        1941. Pteridium aquilinum ssp. typicum
        Western brackenfern Native,
        N.Y. excluded
        NYFA-XCLD
        USDA-NN

        ARS 
        FNA Trop.
        BNA


        Images, wsp

        Family Pteridaceae

        The Pteridaceae E.D.M.Kirchn. (1831), (brakefern family)...

        Subfamily Cryptogrammoideae

        Cryptogramma

        Cryptogramma stelleri
        slender cliffbrake, Steller’s cliffbrake

          Polypodiales — Pteridaceae — Cryptogrammoideae — Cryptogramma
          Cryptogramma Rock-brake N.Y. Status Images Distribution NY NPT
          (S.G.Gmel.) Prantl

          1768. Pteris stelleriS.G.Gmel.
          1882. Cryptogramma stelleriPrantl
          Fragile rockbrake,
          Slender cliff brake,
          Slender rock-brake,
          Steller's rock brake
          Cryptogramme de steller
          Native, C:9,
          Vulnerable,
          DEC-D[1]

          FACU

          Perennial,
          Herb-fern,
          2n=60
          NYFA-3-4
          USDA-N
          NPT Can
          ARS ITIS
          FNA Trop.
          NSE 
          IPN 

          Images, wsp

          Subfamily Pteridoideae

          Pteris

          Pteris multifida
          spider brake
          Pteris multifida, the spider brake was listed as present in New York by the 1955 American Fern Journal, which cited Willard Nelson Clute:
          New York: It is fond of growing on old walls as far north as New York City — Clute (1938, p. 102).[1]

          USDA-NRCS Plants Database lists the 1958 American Fern Journal as the source for the plant being listed in New York, but it doesn't appear to be listed in that volume.[2] Pteris multifida is now been listed as "excluded" in the New York Flora Atlas.


          Polypodiales — Pteridaceae — Pteridoideae — Pteris
          Pteris(excluded taxa) Brake-fern N.Y. Status Images Distribution NY NPT
          Poir.

          1804. Pteris multifidaPoir.
          1938. Pycnodoria multifida(Poir.) Small
           auct.. Pteris serrulatanon L.f.
          Spider brake,
          Spider fern,
          Chinese brake,
          Huguenot fern,
          Saw-leaved bracken
          Introduced from
          temperate Asia,
          tropical Asia,
          N.Y. excluded
          NYFA-XCLD
          USDA-X0

          ARS ITIS
          FNA Trop.

          IPN 

          Images, wsp

            Subfamily Cheilanthoideae

            Pellaea

            Polypodiales — Pteridaceae — Cheilanthoideae — Pellaea
            Pellaea Cliff-brake N.Y. Status Images Distribution NY NPT
            (L.) Link

            Pteris atropurpurea
            Purple cliffbrake,
            Purple-stemmed cliff-brake
            Native,
            Vulnerable,
            DEC-D[1]
            NYFA-4
            USDA-NN
            NPT 





            Images, wsp
            Mett. ex Kuhn
            ssp. glabella

            1858. Pellaea rigida var. rigida
            1869. Pellaea glabella var. glabella
            Pellaea glabella var. bushii
            Pellaea glabella ssp. glabella
            1902. Pellaea atropurpurea var. bushii
            1938. Pellaea atropurpurea fo. glabella
            Smooth cliff-brake
            Slender cliff-brake
            Dwarf cliffbrake
            Simple cliffbrake

            Pelléade glabre
            Pelléade glabre typique
            Pelléade glabre variété glabre
            Native,
            Threatened,
            DEC-B[2]
            NYFA-2
            USDA-NN
            NPT Can

            FNA Trop.



            Images, wsp

            Myriopteris

            Polypodiales — Pteridaceae — Cheilanthoideae — Myriopteris
            Myriopteris Lip-fern N.Y. Status Images Distribution NY NPT
            (Michx.) Grusz & Windham

            1803. Nephrodium lanosumMichx.
            1804. Adiantum vestitumSpreng.
            1806. Cheilanthes vestitaSw.
            1859. Cheilanthes lanosa(Michx.) D.C.Eaton
            2013. M. lanosa(Michx.) Grusz & Windham
            Hairy lipfern,
            Wooly lip-fern
            Native,
            Endangered,
            DEC-A[1]
            NYFA-Z
            USDA-N0
            NPT 
            ARS ITIS
            FNA Trop.
            BNA


            Images, wsp

            Subfamily Vittarioideae

            The Vittarioideae subfamily contains the former families Adiantaceae (maidenhair ferns) and Vittariaceae (shoestring ferns).

              Adiantum

              Adiantum pedatum
              maidenhair fern

                Polypodiales — Pteridaceae — Vittarioideae — Adiantum
                Adiantum Maidenhair-fern N.Y. Status Images Distribution NY NPT
                L.

                1753. Adiantum pedatumL.
                Maidenhair fern,
                Northern maidenhair-fern,
                Northern maidenhair,
                Five-finger fern
                Adiante du Canada
                Native,
                Likely secure,
                DEC-D[1]
                NYFA-4
                USDA-NN
                NPT Can
                ARS 
                FNA Trop.



                Images, wsp
                L.

                1753. Adiantum capillus-veneris
                Venus’s hair fern
                Common maidenhair
                Southern maidenhair

                Introduced from
                southern US,
                N. America native,
                Not naturalized
                NYFA-Xm
                USDA-NN
                NEW-0
                ARS 
                Trop.



                Images, wsp

                Vittaria

                Vittaria appalachiana
                Appalachian shoestring fern
                The genus Vittaria is represented in New York by the endangered Appalachian shoestring fern, which has been found in only a few counties and only in its gametophyte stage.[1]
                1. New York Flora Atlas: Vittaria appalachiana appears only as "minute moss-like gametophytes" in New York.
                Polypodiales — Pteridaceae — Vittarioideae — Vittaria
                Vittaria Shoestring-fern N.Y. Status Images Distribution NY NPT
                Farrar & Mickel (1991)

                Appalachian shoestring fern,
                Appalachian vittaria,
                Appalachian gametophyte fern
                Native,
                Endangered,
                DEC-A[1]
                NYFA-1
                USDA-N0






                Images, wsp

                Family Aspleniaceae

                The classification used here follows Christenhusz & Chase (2014),[1] who place all of eupolypods II in Aspleniaceae Newman (1840).
                1. Maarten J.M. Christenhusz & Mark W. Chase (2014). "Trends and concepts in fern classification". Annals of Botany 113 (9): 571–594. doi:10.1093/aob/mct299.

                Subfamily Cystopteridoideae

                The Cystopteridoideae Ching & Z.R.Wang (1997) (bladder-ferns)...

                Cystopteris

                Cystopteris bulbifera
                bulblet fern

                  Polypodiales — Aspleniaceae — Cystopteridoideae — Cystopteris
                  Cystopteris Bladder-fern N.Y. Status Images Distribution NY NPT
                  (L.) Bernh.

                  1753. Polypodium bulbiferumL.
                  1806. Cystopteris bulbifera(L.) Bernh.
                  1900. Filix bulbifera(L.) Underw.
                  Bulblet fern,
                  Bulblet bladder-fern,
                  Bulblet fragile-fern,
                  Berry fern
                  Cystoptère bulbifère,
                  Cystoptèride bulbifère
                  Native, C:8,
                  Secure,
                  DEC-D[1]

                  Perennial,
                  Herb-fern
                  NYFA-5
                  USDA-NN
                  NPT Can
                  ARS ITIS
                  FNA 



                  Images, wsp
                  (L.) Bernh.

                  1753. Polypodium fragileL.
                  1806. Cystopteris fragilis(L.) Bernh.
                  1848. Cystopteris dickieana R.Sim
                  1900. Filix fragilis(L.) Underw.
                  Brittle bladder-fern,
                  Fragile fern
                  Native, C:7,
                  Likely secure,
                  DEC-D[1]

                  Perennial,
                  Herb-fern
                  NYFA-4
                  USDA-NN
                  NPT 





                  Images, wsp
                  (Michx.) Desv.

                  Cystopteris protrusa ×
                  "Cystopteris hemifragilis"†

                  1803. Nephrodium tenueMichx.
                  1827. Cystopteris tenuis(Michx.) Desv.
                  1889. C. fragilis var. mackayiG.Lawson
                  Mackay's bladder-fern,
                  Mackay's fragile fern,
                  Upland brittle bladder-fern
                  Cystoptère ténue
                  Native, C:7,
                  Likely secure,
                  DEC-D[1]

                  Perennial,
                  Herb-fern
                  NYFA-4
                  USDA-NN
                  NPT 





                  Images, wsp
                  (Weath.) Blasdell

                  1935. C. fragilis var. protrusaWeath.
                  1963. Cystopteris protrusaBlasdell
                  Lowland bladder-fern,
                  Lowland brittle fern,
                  Lowland fragile fern,
                  Southern bladder-fern
                  Native, C:10,
                  Endangered,
                  DEC-A[2]

                  Perennial,
                  Herb-fern
                  NYFA-1
                  USDA-NN
                  NPT 

                  FNA 



                  Images, wsp
                  Cystopteris(excluded taxa) Bladder-fern N.Y. Status Images Distribution NY NPT
                  Cystopteris bulbifera × fragilis

                  Cystopteris bulbifera ×
                  Cystopteris fragilis

                  Hybrid of
                  bulblet bladder-fern &
                  fragile fern
                  Native,
                  N.Y. excluded

                  Perennial,
                  Herb-fern
                  NYFA-XCLD






                  Images, wsp

                  Gymnocarpium

                  Polypodiales — Aspleniaceae — Cystopteridoideae — Gymnocarpium
                  Gymnocarpium Oak-fern N.Y. Status Images Distribution NY NPT
                  (L.) Newman

                  1753. Polypodium dryopterisL.
                  1851. G. dryopteris(L.) Newman
                  1852. Phegopteris dryopteris(L.) Fée
                  1905. Dryopteris linnaeanaC. Chr.
                  1909. Dryopteris dryopteris(L.) Christ
                  1917. Thelypteris dryopteris(L.) Sloss.
                  1966. Carpogymnia dryopterisÁ.& D.Löve
                  Northern oak fern,
                  Common oak fern,
                  Western oak fern
                  Gymnocarpe
                  fougère-du-chêne
                  Native, C:8,
                  Likely secure,
                  DEC-D[1]
                  NYFA-4-5
                  USDA-NN
                  NPT Can
                  ARS ITIS
                  FNA 



                  Images, wsp
                  (Sarvela) Pryer & Haufler

                  Gymnocarpium disjunctum ×
                  Gymnocarpium dryopteris

                  1980. G. dryopteris nssp. brittonianum
                  1993. G. × brittonianumPryer & Haufler
                  Britton's fern,
                  Britton's oak-fern
                  Gymnocarpe de Britton
                  Native,
                  Unranked
                  NYFA-U
                  USDA-NN
                  Can
                  ITIS
                  FNA 



                  Images, wsp
                  Gymnocarpium(excluded) Oak-fern N.Y. Status Images Distribution NY NPT
                  (Rupr.) Ching

                  1845. Polypodium dryopteris var. disjunctum
                  1965. G. disjunctum(Rupr.) Ching
                  Pacific oakfern,
                  Western oak fern
                  Gymnocarpe disjoint,
                  Dryoptère disjointe,
                  Dryoptèride du chêne
                  N. America native,
                  N.Y. excluded
                  NYFA-XCLD
                  USDA-NN
                  Can
                  ITIS
                  FNA Trop.



                  Images, wsp
                  (Hoffm.) Newman

                  1795. Polypodium robertianumHoffm.
                  1850. Phegopteris robertianum(Hoffm.) Fée
                  1851. G. robertianum(Hoffm.) Newman
                  1905. Dryopteris robertiana(Hoffm.) C.Chr.
                  1917. Thelypteris robertiana
                  (Hoffm.) Sloss. ex Rydb.
                  1962. G. dryopteris var. pumilum
                  Scented oakfern,
                  Limestone fern,
                  Robert's oak fern,
                  Northern oak fern
                  Gymnocarpe de Robert
                  N. America native,
                  N.Y. excluded
                  NYFA-XCLD
                  USDA-NN
                  Can
                  ITIS
                  FNA Trop.



                  Images, wsp

                  Subfamily Asplenioideae

                  The Asplenioideae Link (1841) (spleenworts)...

                  Asplenium

                  Asplenium platyneuron
                  ebony spleenwort

                    Polypodiales — Aspleniaceae — Asplenioideae — Asplenium
                    Asplenium Spleenwort N.Y. Status Images Distribution NY NPT
                    var. platyneuron

                    Ebony spleenwort Native, C:5,
                    Secure,
                    DEC-D[1]
                    NYFA-5
                    USDA-NN
                    NPT 





                    Images, wsp
                    L.

                    1753. Asplenium rhizophyllumL.
                    1833. Camptosorus rhizophyllus(L.) Link
                    Walking fern,
                    Walking-fern spleenwort,
                    American walking fern
                    Doradille ambulante,
                    Camptosore à
                    feuilles radicantes
                    Native, C:10,
                    Likely secure,
                    DEC-D[1]
                    NYFA-4-5
                    USDA-NN
                    NPT Can
                    ARS ITIS




                    Images, wsp
                    L.
                    ssp. trichomanes

                    Maidenhair spleenwort Native, C:9,
                    Likely secure,
                    DEC-D[1]
                    NYFA-4
                    USDA-NN
                    NPT 





                    Images, wsp
                    L.
                    ssp. quadrivalensD.E.Mey.

                    Maidenhair spleenwort Native, C:9,
                    Likely secure,
                    DEC-D[1]
                    NYFA-4
                    USDA-NN
                    NPT 





                    Images, wsp
                    L.
                    nssp. lusaticum (D.E.Mey.) Lawalrèe

                    A. t. ssp. trichomanes ×
                    A. t. ssp. quadrivalens

                    1958. Asplenium × lusaticumD.E.Mey.
                    1978. A. trichomanes nssp lusaticum
                    Hybrid
                    Maidenhair spleenwort
                    Native,
                    Unranked,
                    No NY reports

                    NPT 

                    Trop.



                    Images, wsp
                    L.

                    1753. Asplenium ruta murariaL.
                    1928. Asplenium cryptolepisFernald
                    1928. A. cryptolepis var. ohionisFernald
                    1935. A. ruta-muraria var. cryptolepis
                    C. Chr. ex Massey
                    1939. A. ruta-muraria ssp. cryptolepis
                    R.T. Clausen & Wahl
                    Wallrue,
                    Wall-rue spleenwort
                    Doradille des murailles,
                    Mauerraute
                    Native, C:10,
                    Likely secure,
                    DEC-D[1]
                    NYFA-4
                    USDA-NN
                    NPT Can
                    ARS ITIS
                    FNA Trop.



                    Images, wsp

                    Mountain spleenwort Native, C:10,
                    Threatened,
                    DEC-B[2]
                    NYFA-2-3
                    USDA-N0
                    NPT 





                    Images, wsp
                    L.
                    var. americanum
                    (Fernald) Kartesz & Gandhi

                    1793. Scolopendrium vulgareSmall
                    1935. Phyllitis scolopendrium
                    var. americanaFernald
                    1954. Phyllitis fernaldianaÁ. Löve
                    1973. Phyllitis japonica ssp. americana
                    1991. A. scolopendrium var. americanum
                    American
                    hart's-tongue fern,
                    Hart's-tongue fern,
                    Hart's tonguefern
                    Native, C:9,
                    Threatened,
                    DEC-B[2]
                    NYFA-2
                    USDA-N

                    ARS 
                    FNA Trop.



                    Images, wsp
                    Huds.

                    1753. Asplenium ramosumL.
                    1753. A. trichomanes-ramosumL.
                    1762. Asplenium virideHuds.
                    Green spleenwort,
                    Bright-green spleenwort
                    Native, C:10,
                    Endangered,
                    DEC-A[3]
                    NYFA-1
                    USDA-NN
                    NPT 

                    Trop.



                    Images, wsp
                    D.C.Eaton

                    Asplenium montanum ×
                    Asplenium platyneuron

                    1873. Asplenium bradleyiD.C.Eaton
                    1925. Asplenium stotleriWherry
                    1931. Chamaefilix bradleyiFarw.
                    Bradley's spleenwort Native, C:10,
                    No recent reports,
                    Endangered,
                    DEC-A[3]
                    NYFA-Z
                    USDA-N0

                    ITIS
                    FNA 



                    Images, wsp
                    R.R.Scott

                    Asplenium rhizophyllum ×
                    Asplenium platyneuron

                    1865. Asplenium ebenoidesR.R.Scott
                    1937. × Asplenosorus ebenoidesWherry
                    Scott's spleenwort,
                    Dragon tail fern,
                    Walking spleenwort
                    Native,
                    No recent reports
                    NYFA-Z
                    USDA-N0
                    NPT 





                    Images, wsp
                    Asplenium(excluded taxa) Spleenwort N.Y. Status Images Distribution NY NPT
                    L.
                    var. scolopendrium

                    1753. Asplenium scolopendriumL.
                    1844. Phyllitis scolopendriumNewman
                    1881. Scolopendrium
                    scolopendrium
                    H.Karst.
                    Hart's-tongue fern Introduced from
                    Eurasia,
                    northern Africa,
                    N.Y. excluded,
                    Cultivated
                    NYFA-XCLD

                    ARS 
                    Trop.



                    Images, wsp
                    Syme

                    Asplenium ruta-muraria ×
                    Asplenium trichomanes

                    1886. Asplenium clermontaeSyme
                    Hybrid
                    Wallrue &
                    Maidenhair spleenwort
                    Native,
                    Unranked,
                    No NY reports
                    NYFA-D
                    USDA-N0
                    NPT 





                    Images, wsp

                    Subfamily Thelypteridoideae

                    The Thelypteridoideae C.F.Reed (1968) (marsh ferns)

                    Phegopteris

                    Polypodiales — Aspleniaceae — Thelypteridoideae — Phegopteris
                    Phegopteris Beech-fern N.Y. Status Images Distribution NY NPT
                    (Michx.) Watt

                    1753. Polypodium phegopterisL.
                    1803. Polypodium connectileMichx.
                    1826. Lastrea phegopteris(L.) Bory
                    1852. Phegopteris polypodioidesFée
                    1866. Phegopteris connectilis
                    (Michx.) Watt
                    1905. Dryopteris phegopteris(L.) C.Chr.
                    1917. Thelypteris phegopteris(L.) Sloss.
                    Northern beech fern,
                    Narrow beech fern,
                    Long beech fern
                    Phégoptère à
                    segments joint,
                    Fougère du hêtre
                    Native, C:7,
                    Secure,
                    DEC-D[1]

                    FACU

                    Perennial,
                    Fern
                    NYFA-5
                    USDA-NN
                    NPT Can
                    ARS ITIS
                    FNA Trop.



                    Images, wsp
                    (Michx.) Fée

                    1803. Polypodium hexagonopterum
                    Michx.
                    1852. Phegopteris hexagonoptera
                    (Michx.) Fée
                    1905. Dryopteris hexagonoptera
                    (Michx.) C.Chr.
                    1919. Thelypteris hexagonoptera
                    (Michx.) Weath.
                    Broad beech fern,
                    Broad beechfern,
                    Southern beech fern
                    Phégoptère à hexagones,
                    Phégoptéride à hexagones,
                    Thélyptère à hexagones
                    Native,
                    Likely secure,
                    DEC-D[1]
                    NYFA-4-5
                    USDA-NN
                    NPT Can
                    ARS 
                    FNA Trop.

                    IPN LBJ

                    Images, wsp

                    Thelypteris

                    Thelypteris palustris
                    marsh fern
                    As with the hay-scented fern (Dennstaedtia punctilobula) and bracken fern (Pteridium aquilinum), listed above, the New York fern (Thelypteris noveboracensis) is considered to be an "interfering" plant in New York forests. New York ferns produce dense stands that prevent the establishment of tree seedlings.[1] Despite being considered weeds (or conceivably because they are), New York ferns, as well as the other two Thelypteris species, are listed as "exploitably vulnerable" by the New York State Department of Environmental Conservation.[2]
                    Polypodiales — Aspleniaceae — Thelypteridoideae — Thelypteris
                    Thelypteris Maiden-fern N.Y. Status Images Distribution NY NPT
                    Schott
                    var. pubescens(G.Lawson) Fernald

                    1836. Dryopteris thelypteris
                    var. pubescens
                    1864. Lastrea thelypteris var. pubescens
                    1864. T. confluens var. pubescens
                    1910. Dryopteris tremulaChrist
                    1929. T. palustris var. haleana
                    1929. T. palustris var. pubescens
                    Marsh fern,
                    Eastern marsh fern,
                    Northern marsh fern
                    Thélyptère des marais
                    d'Amérique
                    Native, C:4,
                    Secure,
                    DEC-D[1]

                    FACW

                    Perennial,
                    Herb-fern
                    NYFA-5
                    USDA-NN
                    NPT Can
                    ARS ITIS
                    FNA Trop.



                    Images, wsp
                    (L.) Nieuwl.

                    1753. Polypodium noveboracenseL.
                    1803. Nephrodium thelypterioidesMichx.
                    1827. Nephrodium novaeboracenseDesv.
                    1848. Dryopteris noveboracensis
                    1910. Thelypteris noveboracensis
                    1963. Parathelypteris noveboracensis
                    Ching
                    1972. Thelypteris thelypterioidesHolub
                    New York fern
                    Fougère de New York,
                    Thélyptéride de New York
                    Native, C:4,
                    Secure,
                    DEC-D[1]

                    FAC

                    Perennial,
                    Herb-fern
                    NYFA-5
                    USDA-NN
                    NPT 
                    ARS 
                    FNA 



                    Images, wsp
                    (Davenp.) Nieuwl.

                    1894. Aspidium simulatumDavenp.
                    1894. Dryopteris simulataDavenp.
                    1904. Filix simulata(Davenp.) Farw.
                    1910. T. simulata(Davenp.) Nieuwl.
                    1976. Parathelypteris simulataHolttum
                    Massachusetts fern,
                    Bog fern
                    Thélyptère simulatrice
                    Native, C:8,
                    Vulnerable,
                    DEC-D[1]

                    FACW

                    Perennial,
                    Herb-fern
                    NYFA-3-4
                    USDA-N
                    NPT 

                    FNA Trop.



                    Images, wsp
                    Thelypteris(excluded taxa) Maiden-fern N.Y. Status Images Distribution NY NPT
                    Schott
                    var. palustris

                    1834. Thelypteris palustrisSchott
                    Eurasian marsh fern
                    Thélyptère des marais
                    Introduced from
                    Eurasia,
                    N.Y. excluded
                    NYFA-XCLD
                    USDA-






                    Images, wsp

                    Subfamily Woodsioideae

                    The Woodsioideae Fomin (1911) (cliff ferns)...

                    Woodsia

                    Polypodiales — Aspleniaceae — Woodsioideae — Woodsia
                    Woodsia Cliff-fern N.Y. Status Images Distribution NY NPT
                    (L.) R.Br.

                    Acrostichum ilvenseL.
                    Rusty cliff fern,
                    Rusty woodsia
                    Woodsie de l'île d'Elbe
                    Native,
                    Secure,
                    DEC-D[1]
                    NYFA-5
                    USDA-NN
                    NPT 





                    Images, wsp
                    (Spreng.) Torrey
                    ssp. obtusa

                    Bluntlobe cliff fern,
                    Blunt-lobe woodsia
                    Native,
                    Secure,
                    DEC-D[1]
                    NYFA-5
                    USDA-NN
                    NPT 





                    Images, wsp
                    (Bolt.) S.Gray

                    Acrostichum alpinum
                    W. alpina var. bellii
                    W. glabella var. bellii
                    Alpine cliff fern,
                    Alpine woodsia,
                    Northern woodsia
                    Woodsie alpine
                    Native,
                    Endangered,
                    DEC-A[2]
                    NYFA-1
                    USDA-NN
                    NPT 
                    ITIS




                    Images, wsp
                    R.Br. ex Richardson (1823)

                    W. alpina var. glabella
                    W. hyperborea var. glabella
                    Smooth cliff fern,
                    Smooth woodsia
                    Woodsie glabre
                    Native,
                    Endangered,
                    DEC-A[2]
                    NYFA-1
                    USDA-NN
                    glabella/ NPT 

                    FNA 



                    Images, wsp
                    D.C.Eaton (1865)
                    ssp. cathcartiana

                    1908. W. cathcartianaB.L.Rob.
                    W. oregana var. cathcartiana
                    Oregon cliff fern,
                    Oregon woodsia,
                    (tetraploid)
                    Native,
                    Likely extirpated
                    NYFA-Z
                    USDA-NN
                    NPT 
                    ITIS




                    Images, wsp

                    Subfamily Blechnoideae

                    The Blechnoideae ...

                    Onoclea

                    The sensitive fern, so called due to its sensitivity to frost, is very common in areas that are wet for part of the year.

                    Polypodiales — Aspleniaceae — Blechnoideae — Onoclea
                    Onoclea Sensitive fern N.Y. Status Images Distribution NY NPT
                    L.

                    1753. Onoclea sensibilisL.
                    Sensitive fern,
                    Bead fern,
                    Polypod brake,
                    Meadow brake
                    Onoclée sensible
                    Native, C:2,
                    Secure

                    FACW

                    Perennial,
                    Fern
                    NYFA-5
                    USDA-NN
                    NPT 
                    ARS ITIS
                    FNA 
                    BNA


                    Images, wsp

                      Matteuccia

                      With the transfer of Onoclea (above) to Blechnoideae, Christenhusz & Chase (2014) recommended that Matteuccia be moved back to Onoclea.
                        Polypodiales — Aspleniaceae — Blechnoideae — Matteuccia
                        Matteuccia Ostrich fern N.Y. Status Images Distribution NY NPT
                        (L.) Tod.
                        var. pensylvanica(Willd.) C.V.Morton

                        1810. Struthiopteris pensylvanicaWilld.
                        1819. Onoclea pensylvanicaSm.
                        1945. Pteretis pensylvanicaFernald
                        1950. Matteuccia pensylvanicaRaymond
                        1950. Matteuccia struthiopteris
                        var. pensylvanicaC.V.Morton
                        1961. Onoclea struthiopteris
                        var. pensylvanicaB.Boivin
                        Ostrich fern,
                        Shuttlecock fern,
                        Fiddlehead fern
                        Fougère à l'autruche
                        Native, C:5,
                        Likely secure,
                        DEC-D[1]

                        FACW-FAC

                        Perennial,
                        Herb-fern
                        NYFA-4
                        USDA-NN
                        NPT 
                        ARS ITIS
                        FNA 
                        BNA


                        Images, wsp
                        Matteuccia(excluded taxa) Ostrich fern N.Y. Status Images Distribution NY NPT
                        (L.) Tod.
                        var. struthiopteris

                        1753. Osmunda struthiopteris L.
                        1866. Matteuccia struthiopteris(L.) Tod.
                        Ostrich fern,
                        Fougère à l'autruche
                        Introduced from
                        Eurasia,
                        N.Y. excluded
                        NYFA-XCLD
                        USDA-NN
                        NPT 
                        ARS ITIS
                        FNA 
                        BNA


                        Images, wsp

                        Anchistea

                        Polypodiales — Aspleniaceae — Blechnoideae — Anchistea
                        Anchistea Chainfern N.Y. Status Images Distribution NY NPT
                        (L.) C. Presl

                        1771. Blechnum virginicumL.
                        1793. Woodwardia virginica(L.) Sm.
                        1851. Anchistea virginica(L.) C. Presl
                        Virginia chainfern Native, C:9,
                        Secure,
                        DEC-D[1]

                        OBL

                        Perennial,
                        Herb-fern
                        NYFA-5
                        USDA-N
                        NPT 





                        Images, wsp

                        Lorinseria

                        Polypodiales — Aspleniaceae — Blechnoideae — Lorinseria
                        Lorinseria Chainfern N.Y. Status Images Distribution NY NPT
                        (L.) C. Presl

                        1753. Acrostichum areolatumL.
                        1851. Lorinseria areolata(L.) C. Presl
                        1857. Woodwardia areolata(L.) T. Moore
                        Netted chainfern Native, C:10,
                        Vulnerable,
                        DEC-D[1]

                        OBL-FACW

                        Perennial,
                        Herb-fern
                        NYFA-3
                        USDA-N






                        Images, wsp

                        Subfamily Athyrioideae

                        The Athyrioideae (lady-ferns)...

                        Deparia

                        Deparia acrostichoides
                        silvery spleenwort

                          Polypodiales — Aspleniaceae — Athyrioideae — Deparia
                          Deparia False-spleenwort N.Y. Status Images Distribution NY NPT
                          (Sw.) M.Kato

                          Athyrium thelypterioides
                          Diplazium acrostichoides
                          Silvery spleenwort Native, C:8,
                          Secure,
                          DEC-D[1]
                          NYFA-5
                          USDA-NN
                          NPT Can





                          Images, wsp

                          Athyrium

                          Polypodiales — Aspleniaceae — Athyrioideae — Athyrium
                          AthyriumRoth Lady-fern N.Y. Status Images Distribution NY NPT
                          (Willd.) C. Presl

                          1810. Aspidium angustumWilld.
                          1825. Athyrium angustum(Willd.) C.Presl
                          1864. At. filix-femina var. angustum
                          1949. At. filix-femina ssp. angustum
                          Northern lady fern,
                          Common lady fern,
                          Narrow lady fern,
                          Subarctic lady fern
                          Native, C:5,
                          Secure,
                          DEC-D[1]
                          NYFA-5
                          USDA-NN
                          NPT Can
                          ARS ITIS
                          FNA Trop.
                          NSE 
                          IPN 

                          Images, wsp
                          (Michx.) A.A.Eaton

                          1803. Nephrodium asplenioidesMichx.
                          1817. At. asplenioides(Michx.) A.A.Eaton.
                          1923. At. filix-femina var. asplenioides
                          1962. At. filix-femina ssp. asplenioides
                          Southern lady fern,
                          Lowland lady fern
                          Native, C:7,
                          Vulnerable,
                          DEC-D[1]
                          NYFA-3
                          USDA-N0
                          NPT 
                          ITIS
                          FNA Trop.
                          NSE 


                          Images, wsp
                          Athyrium(excluded taxa) Lady-fern N.Y. Status Images Distribution NY NPT
                          (L.) Roth ex Mert.
                          = Athyrium angustum ×
                          Athyrium asplenioides

                          Lady fern,
                          Hybrid of
                          Northern lady fern &
                          Southern lady fern
                          Native,
                          N.Y. excluded
                          NYFA-XCLD



                          BNA


                          Images, wsp
                          (Mett.) Hance

                          1865. Asplenium nipponicumMett.
                          1873. Athyrium nipponicum(Mett.) Hance
                          2011. Anisocampium niponicum
                          Yea C.Liu, W.L.Chiou & M.Kato(Mett.)
                          [2]
                          Japanese painted lady fern,
                          Japanese painted fern
                          Introduced from
                          eastern Asia,
                          No specimens
                          NYFA-0
                          NPT 
                          ARS 
                          Trop.



                          Images, wsp
                          1. 1 2 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.
                          2. [ Liu & al. (2011) "Molecular phylogeny and taxonomy of the fern genus Anisocampium (Athyriaceae)". TAXON 60 (3), June 2011: 824–830.]

                          Family Diplaziopsidaceae

                          The glade fern family includes the genera Diplaziopsis C. Chr. (2–4 spp.) and Homalosorus Pic. Serm. (1 sp.)

                          Homalosorus

                          Homalosorus pycnocarpos
                          narrow-leaved glade fern

                            Polypodiales — Aspleniaceae — Athyrioideae — Homalosorus
                            HomalosorusSw. Twin-sorus fern N.Y. Status Images Distribution NY NPT
                            Serm.

                            1803. Asplenium angustifoliumMichx.
                            nom. illeg., hom. non Jacq. 1786
                            1804. Asplenium pycnocarponSpreng.
                            1906. Athyrium pycnocarponTidestr.
                            1938. Diplazium pycnocarponM.Broun
                            1977. Homalosorus pycnocarposSerm.
                            Glade fern,
                            Narrow-leaved glade fern,
                            Narrow-leaved-spleenwort
                            Athyrie à sores denses,
                            Diplazie à sores denses
                            Native, C:9,
                            Likely secure,
                            DEC-D[1]

                            FAC

                            Perennial,
                            Fern
                            NYFA-4
                            USDA-NN
                            NPT Can
                            ITIS
                            FNA Trop.



                            Images, wsp

                            Family Polypodiaceae

                            The Polypodiaceae J.Presl & C.Presl (1822) (polypody family)

                            Subfamily Dryopteridoideae

                            The Dryopteridoideae Link (1841) (woodferns)...

                            Cyrtomium

                            Polypodiales — Polypodiaceae — Dryopteridoideae — Cyrtomium
                            Cyrtomium Netvein-hollyfern N.Y. Status Images Distribution NY NPT
                            (L.f.) C.Presl

                            1782. Polypodium falcatumL.f.
                            1783. Polypodium japonicumHoutt.
                            1801. Aspidium falcatum(L.f.) Sw.
                            1836. Cyrtomium falcatum(L.f.) C. Presl
                            1891. Dryopteris falcata(L.f.) Kuntze
                            1899. Polystichum falcatum(L.f.) Diels
                            1947. Phanerophlebia falcata(L.f.) Copel.
                            Japanese holly fern,
                            Japanese netvein hollyfern
                            Introduced from
                            Asia,
                            N.Y. excluded
                            NYFA-XCLD
                            USDA-X

                            ARS 
                            Trop.



                            Images, wsp

                              Dryopteris

                              Most Dryopteris (wood fern) species are characterized by sori located near the mid leaf vein. Dryopteris hybrids are very common.
                                Dryopteris diploid species
                                These species have 2n:82 chromosmes (twice the Dryopteris base chromosome number of n:41). D. intermedia and D. Goldiana have been shown to be part of a North American "reticulate complex", in which, along with three other Dryopteris species, they are parents to most of the allopolyploid Dryopteris species shown below.[1][2]
                                Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris
                                Dryopteris(diploid) Wood fern N.Y. Status Images Distribution NY NPT
                                (L.) A.Gray

                                1753. Polypodium marginaleL.
                                1848. Dryopteris marginalisA.Gray
                                Marginal wood fern Native,
                                Secure,
                                DEC-D[1]

                                Perennial,
                                Herb-fern,
                                Anemochore
                                NYFA-5
                                USDA-NN
                                NPT 

                                FNA 



                                Images, wsp
                                (Muhl. ex Willd.) A. Gray

                                1810. Aspidium intermediumMuhl. ex Willd.
                                1920. Thelypteris intermedia(Muhl. ex Willd.) House(?)
                                1825. Nephrodium intermedium(Muhl. ex Willd.) C. Presl
                                1848. Dryopteris intermedia(Muhl. ex Willd.) A. Gray
                                1893. D. spinulosa var. intermedia
                                1950. D. austriaca var. intermedia
                                Evergreen woodfern,
                                Fancy wood fern,
                                Common wood fern
                                Native,
                                Secure,
                                DEC-D[1]

                                Perennial,
                                Herb-fern,
                                Anemochore
                                NYFA-5
                                USDA-NN
                                NPT 

                                FNA 



                                Images, wsp
                                (Hook. ex Goldie) A.Gray

                                1822. Aspidium goldieanumHook. ex Goldie
                                1848. Dryopteris goldieanaA.Gray
                                1910. Thelypteris goldieanaNieuwl.
                                1916. Filix goldianaFarw.
                                1931. Filix-mas goldianaFarw.
                                Goldie's woodfern,
                                Goldie's shield fern,
                                Giant woodfern
                                Dryoptère de Goldie,
                                Dryoptéride de Goldie,
                                Dryoptéris géant
                                Native,
                                Likely secure,
                                DEC-D[1]

                                Perennial,
                                Herb-fern,
                                Anemochore
                                NYFA-4
                                USDA-NN
                                NPT Can
                                ARS ITIS
                                FNA Trop.
                                NSE 
                                IPN 

                                Images, wsp
                                (L.) Schott

                                Fragrant cliff wood-fern,
                                Fragrant woodfern
                                Native,
                                Endangered,
                                DEC-A[2]

                                Perennial,
                                Herb-fern,
                                Anemochore
                                NYFA-1
                                USDA-NN
                                NPT 

                                FNA 



                                Images, wsp
                                Dryopteris allopolyploid species
                                These are fertile species of hybrid origin. Most are allotetraploid, derived from diploid Dryopteris parents, including two of the New York diploid species, D. intermedia and D. goldiana, shown above. The other diploid parents, D. expansa and D. ludoviciana, are North American, but not New York natives. A fifth, likely extinct, diploid parent "D. semicristata" has been hypothesized to form the well-supported model used here. The species Dryopteris clintoniana is allohexaploid, resulting from the diploid D. goldiana (above) and the allotetraploid D. Cristata, below. The parentage of the introduced non-NY-native, Dryopteris filix-mas has not been resolved, but it is probably not part of the same North American "reticulate complex" as the rest.[1]
                                Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris
                                Dryopteris(allopolyploid) Woodfern N.Y. Status Images Distribution NY NPT
                                (L.) A.Gray

                                ♀ D. "semicristata" (SS) ×
                                ♂ D. ludoviciana (LL) → SSLL

                                1753. Polypodium cristatumL.
                                1848. Dryopteris cristataA.Gray
                                Crested shield-fern,
                                Crested woodfern
                                Native,
                                Secure,
                                DEC-D[1]
                                NYFA-5
                                USDA-NN
                                NPT 

                                FNA 



                                Images, wsp
                                (Vill.) H.P.Fuchs

                                ♀ D. "semicristata" (SS) ×
                                ♂ D. intermedia (II) → SSII

                                1777. Polypodium spinulosumO.F.Muell.
                                1786. Polypodium carthusianumVill.
                                1867. Dryopteris spinulosaWatt
                                1959. Dryopteris carthusianaH.P.Fuchs
                                Spinulose shield fern,
                                Spinulose wood fern,
                                Toothed wood fern,
                                Narrow buckler fern
                                Dryoptère spinuleuse
                                Native,
                                Secure,
                                DEC-D[1]
                                NYFA-5
                                USDA-N
                                NPT Can
                                ARS 
                                FNA 



                                Images, wsp
                                (D.C.Eaton) Dowell

                                ♀ D. cristata (SSLL) ×
                                ♂ D. goldiana (GG) → SSLLGG

                                1867. Aspidium cristatum
                                var. clintonianumD.C.Eaton
                                1893. Dryopteris cristata
                                var. clintonianaUnderw.
                                1906. Dryopteris clintonianaDowell
                                1920. Thelypteris clintonianaHouse
                                Clinton's wood fern
                                Dryoptère de clinton
                                Native,
                                Secure,
                                DEC-D[1]
                                NYFA-5
                                USDA-NN
                                NPT 

                                FNA Trop.



                                Images, wsp
                                (Kunze) Clarkson

                                ♀ D. intermedia (II) ×
                                ♂ D. expansa (EE) → IIEE

                                1848. Aspidium campylopterumKunze
                                1915. Dryopteris austriacaWoyn.
                                ex Schinz & Thell.
                                1915. D. spinulosa var. americanaFernald
                                1916. D. dilatata var. americanaBenedict
                                1920. Thelypteris dilatata(Hoffm.) House
                                1930. D. campylopteraClarkson
                                Mountain woodfern,
                                Spreading wood fern,
                                Eastern spreading wood fern
                                Dryoptère arquée
                                Native,
                                Likely secure,
                                DEC-D[1]
                                NYFA-4
                                USDA-N
                                NPT Can
                                ARS ITIS
                                FNA Trop.



                                Images, wsp
                                (W.Palmer)
                                Knowlt., Palmer & Pollard ex Small

                                ♀ D. ludoviciana (LL) ×
                                ♂ D. goldiana (GG) → LLGG

                                1899. D. goldiana ssp. celsaW.Palmer
                                1900. Dryopteris celsaKnowlt. &al.
                                1938. Dryopteris atropalustrisSmall
                                1960. Dryopteris wherryiCrane
                                Log fern Native,
                                Endangered,
                                DEC-A[2]
                                NYFA-1
                                USDA-N

                                ARS ITIS
                                FNA Trop.



                                Images, wsp
                                Dryopteris(excluded) Woodfern N.Y. Status Images Distribution NY NPT
                                (L.) Schott

                                1753. Polypodium filix-masL.
                                1801. Nephrodium filix-masRich.
                                1834. Dryopteris filix-masSchott
                                Male fern,
                                Male wood-fern
                                N. America native,
                                N.Y. excluded
                                NYFA-XCLD
                                USDA-NN
                                NPT 
                                ARS 
                                FNA Trop.
                                NSE BNA


                                Images, wsp
                                Dryopteris named hybrids
                                Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris subsect. named hybrids
                                Dryopteris Woodfern N.Y. Status Images Distribution NY NPT
                                Dryopteris × boottii
                                (Tuck.) Underw.

                                D. cristata (SSLL) ×
                                D. intermedia (II) → ILS

                                1843. Aspidium boottiiTuck.
                                1893. Dryopteris boottii(Tuck.) Underw.
                                1910. Thelypteris boottii(Tuck.) Nieuwl.
                                Boott's wood fern,
                                Glandular swamp fern
                                Native,
                                Vulnerable
                                NYFA-3-5
                                USDA-NN
                                NPT 
                                ITIS




                                Images, wsp
                                Dryopteris × triploidea
                                Wherry

                                D. carthusiana (IISS) ×
                                D. intermedia (II) → IIS

                                1930. D. spinulosa var. fructuosaTrudell
                                1950. D. austriaca var. fructuosaC.V.Morton
                                1958. D. intermedia var. fructuosaWherry
                                1960. Dryopteris × triploideaWherry
                                Triploid woodfern Native,
                                Vulnerable
                                NYFA-3
                                USDA-NN

                                ITIS
                                FNA 



                                Images, wsp
                                Dryopteris × uliginosa
                                (A.Braun ex Dowell) Druce

                                D. carthusiana (IISS) ×
                                D. cristata (SSLL) → ILSS

                                Spinulose crested wood fern Native,
                                Vulnerable
                                NYFA-3
                                USDA-NN






                                Images, wsp
                                Dryopteris × slossoniae
                                Wherry ex Lellinger

                                D. cristata (SSLL) ×
                                D. marginalis (MM) → LMS

                                1894. D. cristata × marginalisDavenport
                                1894. Aspidium cristatum × marginale
                                Davenport
                                1904. Nephrodium slossonaeHahne
                                1942. Dryopteris × slossonaeWherry
                                Crested marginal fern,
                                Miss Slosson's wood fern
                                Native,
                                Threatened
                                NYFA-1-3
                                USDA-NN






                                Images, wsp
                                Dryopteris × pittsfordensis
                                Slossen

                                D. carthusiana (IISS) ×
                                D. marginalis (MM) → IMS

                                1904. Dryopteris pittsfordensisSloss.
                                1904. Nephrodium pittsfordensisWoolson
                                Pittsford's wood fern Native,
                                Endangered
                                NYFA-1
                                USDA-NN

                                ITIS
                                Trop.

                                IPN 

                                Images, wsp
                                Dryopteris × burgessii
                                Boivin

                                D. clintoniana (SSLLGG) ×
                                D. marginalis (MM) → GLMS

                                Burgess' shield fern Native,
                                No recent reports
                                NYFA-Z?
                                USDA-NN






                                Images, wsp
                                Dryopteris × benedictii
                                (Farw.) Wherry

                                D. carthusiana (IISS) ×
                                D. clintoniana (SSLLGG) → GILSS

                                Benedict's wood fern Native,
                                No recent reports
                                NYFA-Z?
                                USDA-NN






                                Images, wsp
                                Dryopteris × dowellii
                                (Farw.) Wherry

                                D. clintoniana (SSLLGG) ×
                                D. intermedia (II) → GILS

                                Dowell's wood fern Native,
                                No recent reports
                                NYFA-Z
                                USDA-NN






                                Images, wsp
                                Dryopteris × mickelii

                                D. clintoniana (SSLLGG) ×
                                D. goldiana (GG) → GGLS

                                Wood fern Native,
                                Likely extirpated
                                NYFA-Z
                                USDA-N






                                Images, wsp
                                Dryopteris × neowherryi
                                W.H.Wagner

                                D. goldiana (GG) ×
                                D. marginalis (MM) → GM

                                Wood fern Native,
                                No recent reports
                                NYFA-Z
                                USDA-NN






                                Images, wsp
                                Dryopteris × separabilis
                                Small (pro sp.)

                                D. celsa (LLGG) ×
                                D. intermedia (II) → GIL

                                Glandular log fern Native,
                                No recent reports
                                NYFA-Z
                                USDA-N0






                                Images, wsp
                                Dryopteris × correllii
                                W.H.Wagner

                                D. carthusiana (IISS) ×
                                D. goldiana (GG) → GIS

                                Correll's wood fern,
                                Poyser's wood fern
                                Native,
                                No recent reports
                                NYFA-?
                                USDA-N0






                                Images, wsp
                                Dryopteris × leedsii
                                Wherry

                                D. celsa (LLGG) ×
                                D. marginalis (MM) → GLM

                                Leeds' wood fern Native,
                                No recent reports
                                NYFA-Z
                                USDA-N0
                                NPT 
                                ITIS




                                Images, wsp
                                  Dryopteris unnamed hybrids
                                  Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris subsect. unnamed hybrids
                                  Dryopteris Woodfern N.Y. Status Images Distribution NY NPT
                                  Dryopteris filix-mas × marginalis

                                  Marginal male fern Native,
                                  Threatened
                                  NYFA-1-3






                                  Images, wsp
                                  Dryopteris celsa × goldiana

                                  D. celsa (LLGG) ×
                                  D. goldiana (GG) → GGL

                                  Hybrid wood-fern Native,
                                  No recent reports
                                  NYFA-Z






                                  Images, wsp
                                  Dryopteris intermedia × marginalis

                                  D. intermedia (II) ×
                                  D. marginalis (MM) → IM

                                  Glandular marginal fern Native,
                                  No recent reports
                                  NYFA-Z






                                  Images, wsp
                                  Dryopteris celsa × cristata

                                  D. celsa (LLGG) ×
                                  D. cristata (SSLL) → GLLS

                                  Hybrid wood-fern Native,
                                  No recent reports
                                  NYFA-Z






                                  Images, wsp
                                  Dryopteris carthusiana × celsa

                                  D. carthusiana (IISS) ×
                                  D. celsa (LLGG) → IGLS

                                  Spinulose wood fern Native,
                                  No recent reports
                                  NYFA-Z






                                  Images, wsp
                                  Dryopteris goldiana × intermedia

                                  D. goldiana (GG) ×
                                  D. intermedia (II) → GI

                                  Wood fern Native,
                                  No recent reports
                                  NYFA-Z






                                  Images, wsp
                                  Dryopteris cristata × goldiana

                                  D. cristata (SSLL) ×
                                  D. goldiana (GG) → GLS

                                  Benedict's wood fern Native,
                                  Unranked
                                  NYFA-U






                                  Images, wsp

                                    Polystichum

                                    Polystichum acrostichoides
                                    Christmas fern
                                    Polystichum Roth (1799) goes by the English vernacular names Christmas fern, holy fern, and sword fern.
                                      Polypodiales — Polypodiaceae — Dryopteridoideae — Polystichum
                                      Polystichum Hollyfern N.Y. Status Images Distribution NY NPT
                                      (Michx.) Schott

                                      1803. Nephrodium acrostichoidesMichx.
                                      1834. Polystichum acrostichoides
                                      (Michx.) Schott
                                      Christmas fern,
                                      Dagger fern
                                      Polystic faux-acrostiche
                                      Native,
                                      Secure
                                      NYFA-5
                                      USDA-NN
                                      NPT 
                                      ARS 
                                      FNA Trop.
                                      BNA


                                      Images, wsp
                                      (Spenner) Fée

                                      1825. Aspidium brauniiSpenner
                                      1852. Polystichum brauniiFée
                                      Braun's holly-fern Native,
                                      Likely secure
                                      NYFA-4
                                      USDA-NN
                                      NPT 

                                      Trop.



                                      Images, wsp
                                      Barrington
                                      [ P. acrostichoides × P. braunii ]

                                      1986. Polystichum × potteriBarrington
                                      Hybrid Christmas fern
                                      & Braun's holly-fern
                                      Native,
                                      Unranked
                                      NYFA-U
                                      USDA-NN


                                      Trop.



                                      Images, wsp
                                      (L.) Roth

                                      1753. Polypodium lonchitisL.
                                      1800. Polystichum lonchitisRoth
                                      1801. Aspidium lonchitisSw.
                                      1924. Polystichum mohrioides var. lemmonii
                                      Northern holly-fern
                                      Polystic faux-lonchitis
                                      Native,
                                      Endangered
                                      NYFA-Z
                                      USDA-NN


                                      FNA 



                                      Images, wsp
                                      Cody

                                      Polystichum acrostichoides ×
                                      Polystichum lonchitis

                                      1924. P. lonchitis fo. marginaleMcColl
                                      1966. P. marginale(McColl) Cody
                                      1968. P. × hagenahiiCody
                                      Hagenah's holly fern,
                                      Hagenah's polystichum,
                                      Hybrid Christmas fern
                                      & northern holly-fern
                                      Polystic de Hagenah
                                      Native,
                                      Unranked,
                                      Questionable[1]

                                      USDA-NN

                                      ITIS
                                      FNA 



                                      Images, wsp
                                      1. Flora of North America states that the rare hybrid Polystichum × hagenahii has only been found growing with both parents in Ontario. NYFA does not list it, but USDA-PLANTS does show it in NY as well as PA, listing the NY source: Lellinger, D.B. (1985) A field manual of the ferns and fern-allies of the U.S. and Canada, Smithsonian Institution, Washington, DC.

                                      Subfamily Polypodioideae

                                      Polypodium

                                      Polypodiales — Polypodiaceae — Polypodioideae — Polypodium
                                      Polypodium Polypody N.Y. Status Images Distribution NY NPT
                                      L.

                                      1753. Polypodium virginianumL.
                                      1818. P. vulgare var. virginianumEaton
                                      1840. P. vulgare var. americanumHook.
                                      1962. P. vulgare ssp. virginianumHultén
                                      1977. P. vinlandicumÁ. Löve & D. Löve
                                      Rock polypody,
                                      Rock-cap fern,
                                      Common polypody
                                      Tripes-de-roches,
                                      Polypode de virginie
                                      Native,
                                      Secure,
                                      DEC-D[1]
                                      NYFA-5
                                      USDA-NN
                                      NPT 

                                      FNA 



                                      Images, wsp
                                      Haufler & Windham

                                      1900. P. vulgare var. oreophilumMaxon
                                      1902. P. vulgare var. acuminatumGilbert
                                      1922. P. virginianum fo. acuminatumFernald
                                      1991. P. appalachianumHaufler & Windham
                                      Appalachian rockcap fern,
                                      Appalachian polypody,
                                      Appalachian rock polypody
                                      Polypode des Appalaches
                                      Native,
                                      Secure
                                      NYFA-5
                                      USDA-NN
                                      NPT 
                                      ARS ITIS
                                      FNA Trop.



                                      Images, wsp
                                      Cusick

                                      Polypodium appalachianum ×
                                      Polypodium virginianum

                                      2002. Polypodium × incognitumCusick
                                      Hybrid polypody Native,
                                      Unranked
                                      NYFA-?
                                      NPT 





                                      Images, wsp

                                      Pleopeltis

                                      Pleopeltis polypodioides
                                      resurrection fern
                                      Pleopeltis polypodioides (resurrection fern) has been excluded from the New York Flora Atlas.[1]
                                      1. USDA-NRCS-PLANTS includes New York in the range of Pleopeltis polypodioides ssp. michauxiana, but the reference for this (House, H.D. 1924. Annotated list of the ferns and flowering plants of New York State. New York State Museum Bulletin No. 254) mentions only that the "gray polypody, or tree fern (Polypodium polypodioides) (L.) Watt, has been found on driftwoud on the Staten Island shore, although its natural range is from Virginia southward."
                                      Polypodiales — Polypodiaceae — Polypodioideae — Pleopeltis
                                      Pleopeltis(excluded taxa) Scaly-polypody N.Y. Status Images Distribution NY NPT
                                      (L.) E.G.Andrews & Windham
                                      var. michauxiana
                                      (Weath.) E.G.Andrews & Windham

                                      1939. Polypodium polypodioides
                                      var. michauxianumWeath.
                                      1993. Pleopeltis polypodioides
                                      var. michauxiana
                                      E.G.Andrews & Windham
                                      2014. Pleopeltis michauxiana
                                      (Weath.) Hickey & Sprunt
                                      Resurrection fern,
                                      Gray's polypody
                                      N. America native,
                                      N.Y. excluded
                                      NYFA-XCLD
                                      USDA-N0

                                      ITIS
                                      FNA Trop.



                                      Images, wsp

                                        Pre-polypod
                                        ferns
                                        Flora of New York Polypodiales
                                        Pinales
                                        Table of
                                        contents
                                        Genus
                                        index
                                        Protected species index Invasive species index
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