< Flora of New York

Dipsacales
Flora of New York Lycophytes
Pre-polypod
ferns
Table of
contents
Genus
index
Protected species index Invasive species index
Order Family Subfamily Genus Common names #
Lycopodiales Lycopodiaceae Huperzioideae Huperzia firmoss 5
Lycopodioideae Lycopodium clubmoss 2
Dendrolycopodium tree clubmoss 3
Spinulum interrupted clubmoss 3
Diphasiastrum ground cedar 7
Lycopodielloideae Lycopodiella bog clubmoss 6
Pseudolycopodiella false bog clubmoss 1
Selaginellales Selaginellaceae Selaginella spikemoss 3
Isoetales Isoetaceae Isoetes quillwort 8

Order Lycopodiales

The Lycopodiales ...

Family Lycopodiaceae

The Lycopodiaceae (clubmoss family).[1]
  1. Lycopodiaceae Troy Weldy & David Werier (2013) New York Flora Atlas. New York Flora Association, Albany, New York.

Subfamily Huperzioideae

The Huperzioideae, known as firmosses, have also been treated as a separate family Huperziaceae. Firmosses differ from other Lycopodiaceae in that they grow in clumps, instead of producing creeping, horizontal stems like other family members.[1] Of the three genera in this group, Huperzia is a temperate and actic genus and is the only one found naturally in New York. Of the other two genera, Phlegmariurus is native to tropical and subtropical regions, and Phylloglossum is native to Australia and New Zealand.
Huperzia
Huperzia lucidula, shining firmoss
Commonly known as firmoss or fir-clubmoss, Huperzia (hoo-per-zee-uh) was named for Johann Peter Huperz, an early 19th century German botanist who specialized in ferns.
    Lycopodiales — Lycopodiaceae — Huperzioideae — Huperzia
    HuperziaBernh. (1801) Firmoss N.Y. Status Images Distribution NY NPT
    (Michx.) Trevis.

    1803. Lycopodium lucidulumMichx.
    1875. Huperzia lucidulaTrevis.
    1939. Urostachys lucidulus
    Herter ex Nessel
    1965. Huperzia selago ssp. lucidula
    Shining firmoss,
    Shining clubmoss,
    Shining fir-clubmoss
    Lycopode brillant,
    Huperzie brillant
    Native, C:5,
    Secure
    NYFA-5
    USDA-NN
    NPT Can
    iNat
    ARS ITIS
    FNA Trop.
    NSE 
    IPN 

    Images, wsp
    (Desv.) A.Löve & D.Löve

    1827. Lycopodium selago var. appressum
    1961. H. appressaA.Löve & D.Löve
    1992. H. appalachianaBeitel & Mickel
    Appalachian firmoss,
    Appalachian clubmoss,
    Appalachian fir-clubmoss,
    Mountain firmoss,
    Mountain fir-clubmoss
    Lycopode des Appalaches
    Native, C:9,
    Rare,
    NYNHP: 3[1]
    NYFA-3
    USDA-NN
    NPT Can
    iNat
    ITIS
    FNA 
    NSE 
    IPN 

    Images, wsp
    (L.) Bernh. ex Schrank & Mart.

    1753. Lycopodium selagoL.
    1778. Lycopodium densumLam.
    1805. Plananthus selagoP.Beauv.
    1821. Lycopodium abietiformeGray
    1829. Huperzia selagoSchrank & Mart.
    1923. Urostachys selagoHerter
    1955. Mirmau selagoH.P.Fuchs
    Northern fir-moss,
    Fir clubmoss
    Lycopode sélagine,
    Huperzie sélagine
    Native, C:10,
    Endangered
    NYFA-1
    USDA-NN
    NPT Can
    iNat
    ARS ITIS
    FNA Trop.
    NSE 
    IPN 

    Images, wsp
    (Cusick) Kartesz & Gandhi

    Huperzia lucidula ×
    Huperzia porophila

    1987. Lycopodium × bartleyiCusick
    1991. H. × bartleyiKartesz & Gandhi
    Bartley's clubmoss Native,
    Unranked
    NYFA-1
    USDA-N0

    iNat
    ITIS
    Trop.



    Images, wsp
    Huperzia(excluded taxa) Firmoss N.Y. Status Images Distribution NY NPT
    (F.E.Lloyd & Underw.) Holub

    1900. Lycopodium porophilum
    F.E.Lloyd & Underw.
    1903. L. selago var. porophilum
    1905. L. lucidulum var. porophilum
    1939. Urostachys lucidulus var. porophilus
    1985. Huperzia porophilaHolub
    Rock clubmoss,
    Rock firmoss,
    Lloyd's clubmoss
    Lycopode porophile
    N. America native,
    N.Y. excluded
    NYFA-XCLD
    USDA-NN
    Can
    iNat

    FNA Trop.



    Images, wsp
    (Clute) Kartesz & Gandhi

    1903. L. lucidulum fo. occidentale
    1932. L. lucidulum var. occidentale
    1966. L. selago var. occidentale
    1967. L. lucidulum var. tryonii
    1991. H. occidentalis
    (Clute) Kartesz & Gandhi
    Pacific Clubmoss

    Lycopode de l'Ouest
    N. America native
    Northwest,
    N.Y. excluded

    Perennial,
    Heerb-forb
    NYFA-XCLD
    USDA-NN

    iNat
    ITIS
    FNA Trop.
    NSE BNA


    Images, wsp
    (Abbe) Kartesz & Gandhi

    Huperzia lucidula ×
    Huperzia selago

    1953. Lycopodium × buttersiiAbbe
    1991. H. × buttersiiKartesz & Gandhi
    Butters' club-moss,
    Butters' firmoss
    Lycopode de Butters
    N. America native,
    N.Y. excluded
    NYFA-XCLD
    USDA-NN
    NPT Can
    iNat
    ITIS
    FNA Trop.
    NSE 


    Images, wsp
    A.Haines

    Huperzia appressa ×
    Huperzia lucidula

    2003. Huperzia × protoporophilaA.Haines
    Hybrid rock firmoss N. America native,
    No N.Y. reports
    NYFA-D
    NPT Can
    iNat

    Trop.

    IPN 

    Images, wsp
    A. Haines

    Huperzia appressa ×
    Huperzia selago

    2003. Huperzia × beiteliiA.Haines illeg.
    2003. Huperzia × josephbeiteliiA. Haines
    Beitel's firmoss,
    Joseph Beitel's clubmoss
    N. America native,
    No N.Y. reports
    NYFA-D
    USDA-NN
    NPT Can
    iNat

    Trop.

    IPN 

    Images, wsp

    Subfamily Lycopodioideae

    Lycopodium
    Running clubmoss
    The genus name Lycopodium (lī-kə-ˈpō-dē-əm) was derived from the Greek lukos (wolf) and podion (small foot). It is commonly known as running clubmoss.

    Also treated as Lycopodium sect. Lycopodium.


      Lycopodiales — Lycopodiaceae — Lycopodioideae — Lycopodium
      Lycopodium Running clubmoss N.Y. Status Images Distribution NY NPT
      L.

      1753. Lycopodium clavatumL.
      L. clavatum var. laurentianum
      L. clavatum var. subremotum
      L. clavatum var. tristachyum
      Common clubmoss,
      Running clubmoss,
      Running pine,
      Running cedar,
      Staghorn clubmoss,
      Elk moss
      Lycopode à massue
      Native, C:5,
      Secure
      NYFA-5
      USDA-NN
      NPT 
      ARS ITIS
      FNA 



      Images, wsp
      (Laest. ex Hartm.) Zinserl. ex Kuzen

      1953. Lycopodium lagopusZinserl. ex Kuzen
      L. clavatum var. brevispicatum
      L. clavatum var. integerrimum
      L. clavatum var. megastachyon
      L. clavatum var. monostachyon
      One-cone clubmoss,
      One-cone ground-pine,
      Running pine,
      Running cedar,
      Arctic stag's-horn clubmoss
      Lycopode patte-de-lapin
      Native, C:4,
      Likely secure
      NYFA-4
      USDA-NN
      NPT 
      ITIS
      FNA 



      Images, wsp
        Dendrolycopodium
        Tree clubmoss
        From Greek δενδρος (denros: tree), λυκος (lukos: wolf), and ποδιων (podion: legs), Dendrolycopodium (den-drō-lī-kə-ˈpō-dē-əm) contains tree-clubmoss species, which are fairly erect and have branched stems. This genus, also treated as Lycopodium sect. Obscura, is characterized by stalked terminal strobili, which contain their sporangia.
          Lycopodiales — Lycopodiaceae — Lycopodioideae — Dendrolycopodium
          Dendrolycopodium Tree clubmoss N.Y. Status Images Distribution NY NPT
          (L.) A.Haines

          1753. Lycopodium obscurumL.
          2003. Dendrolycopodium obscurum
          (L.) A.Haines
          Ground pine,
          Princess-pine,
          Eastern tree clubmoss,
          Rare clubmoss,
          Flat-branch tree clubmoss
          Lycopode obscur
          Native, C:5,
          Secure

          FACU
          NYFA-5
          USDA-NN
          NPT 
          ARS ITIS
          FNA 



          Images, wsp
          (Michx.) A.Haines

          1803. Lycopodium dendroideumMichx.
          1890. L. obscurum var. dendroideum
          1916. L. obscurum var. hybridum
          2003. Dendrolycopodium dendroideum
          (Michx.) A. Haines
          Prickly tree clubmoss,
          Northern tree clubmoss,
          Treelike clubmoss,
          Prickle tree club-moss
          Lycopode dendroïde
          Native, C:5,
          Secure

          FACU
          NYFA-5
          USDA-NN
          NPT 
          ARS ITIS
          FNA 



          Images, wsp
          (W.H.Wagner, Beitel, & Moran) A.Haines

          L. obscurum var. isophyllum
          Pennsylvania clubmoss,
          Hickey's tree club-moss
          Native, C:5,
          Secure

          Perennial,
          Herb

          NYFA-5
          USDA-NN
          NPT 
          ITIS
          FNA 



          Images, wsp
            Spinulum

            Spinulum (ˈspīn-yə-ləm) is also treated as Lycopodium sect. Annotina.

            Lycopodiales — Lycopodiaceae — Lycopodioideae — Spinulum
            Spinulum Bristly clubmoss N.Y. Status Images Distribution NY NPT
            (L.) A.Haines

            1753. Lycopodium annotinumL.
            1772. Lycopodium dubiumZoëga
            2003. Spinulum annotinumA.Haines
            Common bristly clubmoss,
            Common stiff clubmoss,
            Interrupted clubmoss
            Lycopode interrompu
            Native, C:5,
            Secure
            NYFA-5
            USDA-NN
            NPT Can
            ARS ITIS
            FNA 



            Images, wsp
            (Nessel) A.Haines

            1772. Lycopodium dubiumauct. non Zoëga
            1842. L. annotinum var. pungensDesv.
            1842. L. pungensLa Pylaie ex Desv.
            1843. L. annotinum var. montanumTuck.
            1940. L. canadenseNessel
            1968. L. annotinum ssp. pungensHultén
            2003. Spinulum canadenseA.Haines
            Northern bristly clubmoss,
            Northern stiff clubmoss,
            Northern interrupted clubmoss
            Native, C:10,
            Endangered
            NYFA-1-2
            NPT Can

            Trop.



            Images, wsp

            Hybrid bristly clubmoss Native,
            Unranked
            NYFA-U






            Images, wsp
              Diphasiastrum

              Also treated as Lycopodium sect. Complanata.

              Lycopodiales — Lycopodiaceae — Lycopodioideae — Diphasiastrum
              DiphasiastrumHolub (1975) Ground-cedar N.Y. Status Images Distribution NY NPT
              (Dill. ex A.Braun) Holub

              1848. Lycopodium digitatumDill. ex A.Braun
              1901. L. complanatum var. flabelliforme
              1911. L. flabelliforme(Fernald) Blanch.
              1965. Diphasium complanatum
              ssp. flabelliforme
              1975. Diphasiastrum digitatum
              (Dill. ex A.Braun) Holub
              Southern groundcedar,
              Southern ground-cedar,
              Fan club-moss,
              Southern running-pine
              Lycopode en éventail,
              Courants verts,
              Lycopode aplati
              variété en éventail
              Native, C:4,
              Secure
              NYFA-5
              USDA-NN
              NPT Can
              ARS ITIS
              FNA Trop.

              IPN 

              Images, wsp
              (Pursh) Holub (1975)

              Lycopodium tristachyum Pursh (1814)'
              Diphasium tristachyum (Pursh) Rothm.'
              L. complanatum chamaecyparissus
              Blue ground-cedar,
              Deep-root clubmoss
              Native, C:4,
              Secure
              NYFA-5
              USDA-NN

              ITIS
              FNA 



              Images, wsp
              (Willd.) Holub (1975)

              Diphasiastrum sitchense ×
              Diphasiastrum tristachyum

              Lycopodium × sabinifoliumWilld.
              Lycopodium sabinifoliumWilld.
              Lycopodium armatumDesv.
              Ground-fir,
              Savinleaf groundpine,
              Savin-leaved groundpine,
              Cypress clubmoss
              Native,
              Vulnerable
              NYFA-3
              USDA-NN

              ARS ITIS
              FNA 



              Images, wsp
              (Rouy) Holub (1975)
              [ complanatum × tristachyum ]

              Lycopodium × zeilleri
              Diphasium zeilleri (Rouy) Damboldt'
              Zeiller's running-pine,
              Zeiller's clubmoss
              Native,
              Threatened
              NYFA-2
              USDA-NN

              ITIS
              FNA 



              Images, wsp
              (L.) Holub

              1753. Lycopodium complanatumL.
              Diphasium complanatum(L.) Rothm.
              Lycopodium ancepsWallr.
              Diphasium anceps(Wallr.) Á,D.Löve
              1975. Diphasiastrum complanatum(L.) Holub
              Northern groundcedar,
              Trailing clubmoss,
              Northern running-pine,
              Lycopode aplati
              Native, C:8,
              Endangered
              NYFA-1
              USDA-NN

              ARS ITIS
              FNA 



              Images, wsp
              (Rupr.) Holub (1975)

              Lycopodium sitchense Rupr. (1845)'
              Diphasium sitchense (Rupr.) Á,D.Löve'
              Lycopodium sabinifolium sitchense
              Sitka groundcedar,
              Alaskan clubmoss,
              Sitka club-moss,
              Lycopode de Sitka
              Native, C:10,
              Endangered
              NYFA-1
              USDA-NN

              ARS ITIS
              FNA 



              Images, wsp
              (House) Holub (1975)

              Diphasiastrum digitatum ×
              Diphasiastrum tristachyum

              Lycopodium × habereriHouse
              L. complanatum var. habereri
              L. tristachyum var. habereri
              Haberer's clubmoss Native,
              Unranked
              NYFA-U
              USDA-NN

              ITIS
              FNA 



              Images, wsp

                Subfamily Lycopodielloideae

                Lycopodiella

                Bog clubmoss. (Lycopodiella in Latin means little wolf's-foot.)

                Lycopodiales — Lycopodiaceae — Lycopodielloideae — Lycopodiella
                LycopodiellaHolub (1964) Bog clubmoss N.Y. Status Images Distribution NY NPT
                (L.) Holub

                1753. Lycopodium inundatumL.
                1805. Plananthus inundatus(L.) P.Beauv.
                1852. Lepidotis inundata(L.) Opiz
                1912. Lepidotis inundata(L.) Börner
                1964. Lycopodiella inundata(L.) Holub
                Northern bog clubmoss,
                Marsh clubmoss,
                Inundated clubmoss
                Lycopodielle inondé
                Native, C:8,
                Secure
                NYFA-5
                USDA-NN
                NPT 
                ARS ITIS
                FNA Trop.



                Images, wsp
                (Chapm.) Cranfill

                1843. Lycopodium inundatum var. bigelovii
                1878. Lycopodium inundatum var. appressum
                1900. Lycopodium appressum
                F.E.Lloyd & Underw.
                1981. Lycopodiella appressa
                (Chapm.) Cranfill
                1985. Lycopodiella bigelowii(Tuck.) Holub
                Southern bog clubmoss,
                Appressed bog-clubmoss,
                Chapman's club-moss
                Native, C:7,
                Likely secure
                NYFA-4
                USDA-NN
                NPT 
                ITIS
                FNA 



                Images, wsp
                (L.) Cranfill

                1753. Lycopodium alopecuroidesL.
                1944. Lepidotis alopecuroides(L.) Rothm.
                1981. Lycopodiella alopecuroides(L.) Cranfill
                Foxtail bog-clubmoss,
                Foxtail clubmoss
                Native, C:7,
                Vulnerable

                NYFA-3
                USDA-N0
                NPT 
                ITIS
                FNA 



                Images, wsp
                (Eiger) Cranfill (1981)

                L. alopecuroides ×
                L. appressa

                Copeland's clubmoss Native,
                Unranked
                NYFA-U
                USDA-N0

                ITIS




                Images, wsp
                A.Haines (2003)

                L. appressa ×
                L. inundata

                Gilman's clubmoss Native,
                Unranked
                NYFA-U

                ITIS




                Images, wsp
                Lycopodiella(excluded taxa) Bog clubmoss N.Y. Status Images Distribution NY NPT
                ( R.J.Eaton) A.Haines (2002)

                L. alopecuroides ×
                L. inundata

                Lycopodium inundatum var. robustum
                Bog clubmoss Native,
                N.Y. excluded
                NYFA-U






                Images, wsp
                  Pseudolycopodiella
                  Lycopodiales — Lycopodiaceae — Lycopodielloideae — Pseudolycopodiella
                  Pseudolycopodiella False bog-clubmoss N.Y. Status Images Distribution NY NPT
                  (L.) Holub

                  1753. Lycopodium carolinianumL.
                  1968. Lycopodiella caroliniana(L.) Pic.Serm.
                  1983. Pseudolycopodiella carolinianaHolub
                  1983. Pseudolycopodiella meridionalisHolub
                  Carolina clubmoss,
                  Slender clubmoss,
                  Slender bog club-moss
                  Native, C:9,
                  Endangered,
                  NYNHP: 1[1]
                  NYFA-1
                  USDA-N0
                  NPT 
                  ARS ITIS
                  FNA Trop.



                  Images, wsp

                  Order Selaginellales

                  Family Selaginellaceae

                  Selaginella

                  Three spikemoss (Selaginella) species have been found growing wild in New York.
                    Selaginellales — Selaginellaceae — Selaginella
                    Selaginella Spikemoss N.Y. Status Images Distribution NY NPT
                    (L.) Spring

                    1753. Lycopodium rupestreL.
                    1838. Selaginella rupestris(L.) Spring
                    Rock spikemoss,
                    Dwarf spikemoss,
                    Northern selaginella
                    Sélaginelle des rochers,
                    Sélaginelle rupestre
                    Native, C:9,
                    Likely secure

                    UPL

                    Perennial,
                    Herb-forb
                    NYFA-4
                    NRCS-NN
                    NPT 
                    iNat
                    ITIS
                    FNA 



                    Images, wsp
                    (L.) Spring

                    1753. Lycopodium apodumL.
                    1840. Selaginella apusSpring
                    1840. Selaginella apoda(L.) Spring
                    Meadow spikemoss,
                    Creeping spikemoss
                    Sélaginelle apode
                    Native, C:6,
                    Threatened,
                    Under review[1]

                    FACW
                    NYFA-2?
                    NRCS-N0
                    NPT 
                    iNat
                    ITIS
                    FNA 



                    Images, wsp
                    W.R.Buck

                    1977. Selaginella eclipesW.R.Buck
                    1996. S. apoda ssp. eclipesSkoda
                    Hidden spikemoss,
                    Buck's meadow spikemoss,
                    Creeping spikemoss
                    Native, C:8,
                    Endangered

                    FACW
                    NYFA-1
                    NRCS-NN
                    NPT 
                    ITIS
                    FNA Trop.



                    Images, wsp
                    Selaginella(excluded taxa) Spikemoss N.Y. Status Images Distribution NY NPT
                    (L.) P. Beauv. ex Mart. & Schrank

                    1753. Lycopodium selaginoidesL.
                    1805. S. spinosaP.Beauv. illeg.
                    1841. S. selaginoides(L.)
                    P.Beauv. ex Mart. & Schrank
                    Club spikemoss
                    Northern spikemoss
                    Northern spike-moss
                    Prickly mountain-moss
                    Lesser-club-moss

                    N. America native,
                    N.Y. excluded

                    FACW

                    Perennial,
                    Herb-forb
                    NYFA-XCLD
                    USDA-NN
                    NPT 
                    ARS 




                    Images, wsp

                    Order Isoetales

                    Family Isoetaceae

                    Isoetes

                    Isoetales — Isoetaceae — Isoetes
                    Isoetes Quillwort N.Y. Status Images Distribution NY NPT
                    A.Braun

                    1846. Isoetes engelmanniiA.Braun
                    1905. Isoetes validaClute
                    1997. Isoetes appalachiana
                    D.F.Brunt. & D.M.Britton
                    Engelmann's quillwort,
                    Appalachian quillwort
                    Isoète d'Engelmann
                    Native, C:6,
                    Secure
                    NYFA-5
                    USDA-NN
                    NPT Can
                    ITIS
                    FNA Trop.
                    NSE BNA
                    IPN 

                    Images, wsp
                    Durieu
                    ssp. muricataA.Löve & D.Löve[1]

                    1789. Isoetes setaceaLam. p.p.
                    1827. Isoetes tenellaLéman ex Desv.
                    1861. Isoetes echinosporaDurieu
                    1864. Isoetes brauniiDurieu
                    1864. Isoetes muricataDurieu
                    1867. I. echinospora var. braunii
                    1961. I. echinospora ssp. muricata
                    Spiny-spore quillwort,
                    Spring quillwort,
                    Braun's quillwort,
                    Bristlelike quillwort
                    Isoète à spores
                    épineuses,
                    Isoète à spores
                    spinuleuses
                    Native, C:7,
                    Likely secure
                    NYFA-4
                    USDA-?
                    NPT Can
                    ARS ITIS
                    FNA Trop.



                    Images, wsp
                    A.Braun ex Engelm.

                    1867. I. tuckermaniiA.Braun ex Engelm.
                    1900. I. tuckermanii var. borealis
                    1981. I. acadiensisKott
                    1991. I. tuckermanii ssp. acadiensis
                    Tuckerman's quillwort
                    Isoète de Tuckerman
                    Native, C:10,
                    Likely secure
                    NYFA-4
                    USDA-NN
                    NPT 
                    ITIS
                    FNA 



                    Images, wsp
                    R.Dodge

                    Isoetes echinospora ×
                    Isoetes engelmannii

                    1896. Isoetes eatoniiR.Dodge
                    1900. I. gravesiiA.A.Eaton
                    I. eatonii var. gravesiiA.A.Eaton
                    Eaton's quillwort Native,
                    Vulnerable

                    NYFA-3
                    USDA-NN

                    ITIS




                    Images, wsp
                    L.

                    1753. Isoetes lacustrisL.
                    1864. Isoetes macrosporaDurieu
                    1900. Isoetes hieroglyphicaA.A.Eaton
                    1922. I. macrospora fo. hieroglyphica
                    Lake quillwort,
                    Carved quillwort,
                    Warty Quillwort
                    Isoète lacustre,
                    Isoète hiéroglyphique
                    Native, C:10,
                    Rare
                    NYFA-3
                    USDA-NN
                    NPT 
                    ITIS
                    FNA 



                    Images, wsp
                    D.F.Brunt.

                     auct. I. riparia var. canadensis (i)
                    1903. I. canadensis var. robbinsii
                    2015. I. septentrionalisD.F. Brunt.
                    Northern shore quillwort,
                    Northern quillwort
                    isoète du Nord
                    Native, C:10,
                    Endangered

                    OBL

                    Perennial,
                    Herb-forb
                    NYFA-1
                    USDA-N0
                    NPT Can
                    ITIS
                    FNA 



                    Images, wsp
                    I. engelmannii × septentrionalis

                    Isoetes engelmannii ×
                    Isoetes septentrionalis

                     auct. Isoetes × brittonii
                     auct. Isoetes engelmannii × riparia
                    Hybrid of
                    Engelmann's quillwort &
                    northern shore quillwort
                    Native,
                    No recent reports

                    NYFA-Z?
                    USDA-N0
                    NPT 
                    ARS ITIS




                    Images, wsp
                    1. The New York Flora Atlas and New England Wild Flower Society list the spiny-spore quillwort as Isoëtes echinospora ssp. muricata because the leaves of the North American plants of bear stomata, while those of the European plants do not. The other sources used here treat ssp. muricata as a synonym.

                    Isoetes (excluded)

                    Isoetales — Isoetaceae — Isoetes
                    Isoetes(excluded taxa) Quillwort N.Y. Status Images Distribution NY NPT
                    Engelm. ex A.Braun[1]

                    1846. I. ripariaEngelm. ex A.Braun
                    18??. I. saccharataEngelm. in A.Gray
                    1891. Calamaria saccharata
                    (Engelm.) Kuntze
                    1901. I. canadensis(Engelm.) A.A.Eaton
                    1949. I. riparia fo. saccharata
                    Shore quillwort,
                    Canada shore quillwort
                    Native,
                    N.Y. excluded
                    NYFA-XCLD
                    USDA-NN
                    NPT 
                    ITIS
                    Trop.
                    BNA


                    Images, wsp
                    A.A.Eaton

                    Isoetes lacustris ×
                    Isoetes tuckermanii

                    1900. Isoetes × heterosporaA.A.Eaton
                    Isoetes tuckermanii var. harveyi
                    Harvey's quillwort Native,
                    N.Y. excluded
                    NYFA-XCLD
                    USDA-NN
                    NPT 
                    ITIS
                    FNA 



                    Images, wsp
                    D.F. Brunt. & D.M. Britton

                    Isoetes septentrionalis ×
                    Isoetes tuckermanii

                    New England quillwort Native,
                    N.Y. excluded
                    NYFA-XCLD
                    USDA-N0
                    NPT 
                    ITIS
                    FNA 



                    Images, wsp
                    1. Isoetes riparia was apparently misapplied to reports of Isoetes septentrionalis in New York. (NYFA (2016)

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