Itaboraian

The Itaboraian (Spanish: Itaboraiense) age is a period within the Early Eocene geologic time (53.0–50.0 Ma) epoch of the Paleogene, used more specifically with South American land mammal ages (SALMA). It follows the Riochican and precedes the Casamayoran age.[1]

Stratigraphy and location of the Itaboraí Basin.
Crocodile skull indicates Sahitisuchus find.

Etymology

This age is named after the Itaboraí Formation in the Itaboraí Basin in Itaboraí, a Greater Rio municipality in the state of Rio de Janeiro in Brazil not very far away from Niterói.

Formations
Locations of Itaboraian formations
Formation
bold is type		 Country Basin Notes
Itaboraí Formation BrazilItaboraí Basin
[2]
Bogotá Formation ColombiaAltiplano Cundiboyacense
[3]
Cerrejón Formation ColombiaCesar-Ranchería Basin
[4]
Chota Formation PeruBagua Basin
[5][6]
Koluel Kaike Formation ArgentinaGolfo San Jorge Basin
[7]
Las Flores Formation ArgentinaGolfo San Jorge Basin
[8]
Maíz Gordo Formation ArgentinaSalta Basin
[9]
Mealla Formation ArgentinaSalta Basin
[10]
Mogollón Formation PeruTalara & Tumbes Basins
[11]
Muñani Formation PeruAltiplano Basin
[12]
Peñas Coloradas Formation ArgentinaGolfo San Jorge Basin
[13]
Río Loro Formation ArgentinaSierras Pampeanas
[14]

Fossils
Group Fossils Formation Notes
MammalsAsmithwoodwardia scotti, Austropediomys marshalli, Bergqvistherium primigenia, Bobbschaefferia fluminensis, Camargomendesia pristina, Carodnia vieirai, Carolocoutoia ferigoloi, Carolopaulacoutoia itaboraiensis, Colbertia magellanica, Derorhynchus singularis, Didelphopsis cabrerai, Eobrasilia coutoi, Epidolops ameghinoi, Gashternia carioca, Gaylordia macrocynodonta, Gaylordia mater, Guggenheimia brasiliensis, Guggenheimia crocheti, Homalostylops atavus, Itaboraidelphys camposi, Lamegoia conodonta, Marmosopsis juradoi, Miguelsoria parayirunhor, Minusculodelphis minimus, M. modicum, Mirandatherium alipioi, Monodelphopsis travassosi, Palaeocladosictis mosesi, Paranisolambda prodromus, Patene simpsoni, Periprotodidelphis bergqvistae, Procaroloameghinia pricei, Protodidelphis mastodontoides, P. vanzolinii, Protolipterna ellipsodontoides, Ricardocifellia protocenica, Riolestes capricornicus, Riostegotherium yanei, Tetragonostylops apthomasi, Victorlemoinea prototypica, Xenodelphis doelloi, Zeusdelphys complicatus, ?Arminiheringia sp., Henricosbornia sp., Australidelphia indet., Borhyaenidae indet., Didelphidae indet., Didolodontidae indet., Herpetotheriidae indet., Microbiotheriidae indet., Notoungulata indet., Paucituberculata indet., Pediomyidae indet., Peradectidae indet., Polydolopidae indet., Protodidelphidae indet.Itaboraí
[15]
[16][17]
[18]
[2][19]
[20]
Ernestokokenia yirunhor, Gashternia ctalehor, Henricosbornia waitehor, Seudenius cteronc, Shecenia ctirneru, ?Isotemnus ctalego, ?Peripantostylops orehor, ?Polydolops kamektsen, Kibenikhoria sp., Archaeopithecidae indet., Borhyaenidae indet., ?Litopterna indet., ?Notostylopidae indet.Koluel Kaike
[21]
Anisolambda sp., Antepithecus sp., Asmithwoodwardia sp., Bardalestes sp., Bobbschaefferia sp., Camargomendesia sp., Carolopaulacoutoia sp., Colbertia sp., Derorhynchus sp., Didelphopsis sp., Epidolops sp., Ernestokokenia sp., Gashternia sp., Guggenheimia crocheti, Henricosbornia sp., Isotemnus sp., Itaboraitherium sp., Kibenikhoria sp., Marmosopsis sp., Mirandatherium sp., Monodelphopsis sp., Peripantostylops sp., Polydolops sp., Procaroloameghinia pricei, Protodidelphis sp., Shecenia sp., Trigonostylops sp., Victorlemoinea sp., Woodburnodon sp., Ceratodontidae indet., ?Patene indet.Las Flores
[22]
Simpsonotus major, S. praecursorMealla
[23][24]
Carodnia inexpectansMogollón
[25][26]
Chulpasia mattaueri, Peradectes austrinum, Perutherium altiplanense, Sillustania quechuense, Umayodus raimondi, Didelphidae indet., ?Didolodontidae indet., Notoungulata sp., Pediomyidae indet., Proteutheria indet.Muñani
[27]
[28][29]
BirdsDiogenornis fragilis, Eutreptodactylus itaboraiensis, Itaboravis elaphrocnemoides, Paleopsilopterus itaboraiensisItaboraí
[19][30]
[31][32]
Latidae indet., Siluriformes indet., Tetragonopterinae indet.Muñani
[29]
Reptiles &
amphibians		Acherontisuchus guajiraensis, Anthracosuchus balrogus, Carbonemys cofrinii, Cerrejonemys wayuunaiki, Cerrejonisuchus improcerus, Pelomedusoides sp., Puentemys mushaisaensis, Titanoboa cerrejonensisCerrejón
[33][34]
?Madtsoia sp.Koluel Kaike
[21]
Notocaiman stromeriPeñas
Coloradas
[35]
Bretesuchus bonaparteiMaíz Gordo
[36]
Apodops pricei, Coniophis cf. precedens, Corallus priscus, Hechtophis austrinus, Hoffstetterella brasiliensis, Itaboraiophis depressus, Madtsoia camposi, Paraungaliophis pricei, Paulacoutophis perplexus, Sahitisuchus fluminensis, Waincophis cameratus, Waincophis pressulus, Xenopus romeri, ? Russellophiidae indet.Itaboraí
[37][38]
Lorosuchus nodosus, Pelomedusoides cf. argentinensisRío Loro
[39]
cf. Coniophis sp., ?Roxochelys vilavilensis, Crocodylia indet., ?Leptodactylidae indet.Muñani
[29]
FishesDipnoi sp., Elopomorpha sp.Cerrejón
[33]
Lepidosiren cf. paradoxa, Ceratodus sp.Muñani
[29]
MolluscsBulimulus fazendicusItaboraí
[40]
FloraMenispina evidensBogotá
[41]
Stephania palaeosudamericanaBogotá, Cerrejón
[42]
Malvaciphyllum macondicus, Petrocardium cerrejonense, Petrocardium wayuuorumCerrejón
[43][44]

References
  1. Paleo Database: Itaboraian
  2. Oliveira & Goin, 2011
  3. Bayona et al., 2012, p.103
  4. Jaramillo et al., 2007, p.168
  5. Chacaltana et al., 2015, p.2
  6. Castro Medina, 2010, p.53
  7. Raigemborn et al., 2010, p.251
  8. Raigemborn et al., 2010, p.249
  9. Narváez, 2009, p.24
  10. Sánchez & Marquillas, 2010, p.389
  11. Quispe Alcalá, 2010, p.21
  12. Gelfo & Goin, 2011
  13. Clyde et al., 2014, p.303
  14. Pol & Powell, 2011, S9
  15. Oliveira et al., 2016
  16. Carneiro et al., 2018
  17. Carneiro, 2018
  18. De Oliveira & Goin, 2015
  19. São José de Itaboraí at Fossilworks.org
  20. São José 700 m at Fossilworks.org
  21. Cañadón Hondo at Fossilworks.org
  22. Las Flores at Fossilworks.org
  23. Road linking Tres Cruces and Mina Aguilar at Fossilworks.org
  24. Between Pisungo and Rio Grande at Fossilworks.org
  25. Antoine et al., 2015
  26. Carodnia inexpectans at Fossilworks.org
  27. Gelfo & Sigé, 2011
  28. Laguna Umayo at Fossilworks.org
  29. LURMU at Fossilworks.org
  30. Diagenornis at Fossilworks.org
  31. Itaboravis at Fossilworks.org
  32. Paleopsilopterus at Fossilworks.org
  33. La Puente Pit at Fossilworks.org
  34. Anthracosuchus at Fossilworks.org
  35. Notocaiman stromeri at Fossilworks.org
  36. Bretesuchus at Fossilworks.org
  37. Itaboraí snakes at Fossilworks.org
  38. Portland Quarry at Fossilworks.org
  39. El Cadillal at Fossilworks.org
  40. Salvador & Simone, 2013, p.7
  41. Menispina evidens at Fossilworks.org
  42. Stephania palaeosudamericana at Fossilworks.org
  43. Cerrejón 0315 at Fossilworks.org
  44. Coalbeds 100 at Fossilworks.org

Bibliography
Itaboraí Formation
Bogotá Formation
Cerrejón Formation
Chota Formation
Las Flores Formation
Koluel Kaike Formation
Maíz Gordo Formation
Mealla Formation
Mogollón Formation
Muñani Formation
Peñas Coloradas Formation
Río Loro Formation
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