Gamerabaena

Gamerabaena is an extinct genus of baenid turtle which existed in North Dakota during the late Cretaceous period. It is known from a single fragmentary skull that was found in the Maastrichtian-age Hell Creek Formation. It contains the species Gamerabaena sonsalla.[1] Gamerabaena is similar to the genus Palatobaena, but it differs in its lack of a posterior expansion of the triturating (or chewing) surface, a somewhat rectangular skull, and a wide angle between the maxillae. Gamerabaena also has a lingual ridge on the inner side of the jaw that is not seen in Palatobaena.[1]

Gamerabaena Temporal range: Late Cretaceous
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Family:	†Baenidae
Genus:	†Gamerabaena Lyson and Joyce, 2010
Species
G. sonsalla Lyson and Joyce, 2010 (type)

Gamerabaena is considered the sister taxon of Palatobaena and shares features with both Palatobaena and Plesiobaena. These features, which include slightly upturned eye sockets, are seen as intermediate between the two other genera. While Gamerabaena is known only from the skull, it may belong to the same species as "Baena" hayi, which is known primarily from the shell.[1]

The genus is named after Gamera, a giant flying, fire-breathing turtle from a series of Japanese tokusatsu films.[1]

Phylogeny

Cladogram after Tyler R. Lyson and Walter G. Joyce (2009).[2]

Pleurosternon bullockii

Glyptops plicatulus

Baenidae

Neurankylinae

	Neurankylus eximius

	Trinitichelys hiatti

Eubaeninae

Hayemys latifrons

Baenodda

Plesiobaenini

Plesiobaena antiqua

Peckemys brinkman

Cedrobaena putorius

Gamerabaena sonsalla

Palatobaena cohen

	Palatobaena bairdi

	Palatobaena gaffneyi

Eubaenini

	Boremys pulchra

	Boremys grandis

	Eubaena cephalica

Goleremys mckennai

Stygiochelys estesi

	Baena arenosa

	Chisternon undatum

References

Lyson, T.R.; Joyce, W,G. (2010). "A new baenid turtle from the Upper Cretaceous (Maastrichtian) Hell Creek Formation of North Dakota and a preliminary taxonomic review of Cretaceous Baenidae" (PDF). Journal of Vertebrate Paleontology. 30 (2): 394–402. doi:10.1080/02724631003618389.CS1 maint: multiple names: authors list (link)
Tyler R. Lyson & Walter G. Joyce (2009). "A Revision of Plesiobaena (Testudines: Baenidae) and an Assessment of Baenid Ecology Across the K/T Boundary". Journal of Paleontology. 83 (6): 833–853. doi:10.1666/09-035.1.

Testudines

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Subclass: Anapsida
Order: Testudines

Suborder

Superfamily

Family

Genus

Cryptodira

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Bouliachelys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Alienochelys †Berruchelus †Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	Chrysemys Clemmys Deirochelys Emys Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Tortoises	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys Psammobates Pyxis Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

†Baenoidea

†Baenidae	†Arvinachelys †Cedrobaena †Chisternon †Denazinemys †Gamerabaena †Neurankylus †Palatobaena †Peckemys †Plesiobaena
†Pleurosternidae	†Dinochelys †Dorsetochelys †Glyptops †Pleurosternon †Selenemys

Chelydridae	†Acherontemys Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Eurysternidae	†Achelonia †Chelonides †Eurysternum †Hydropelta †Idiochelys †Palaeomedusa †Parachelys
†Macrobaenidae	†Anatolemys †Aurorachelys †Kirgizemys †Osteopygis
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Archelon †Atlantochelys †Chelosphargis †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
†Sinemydidae	†Jeholochelys †Liaochelys †Manchurochelys †Ordosemys †Sinemys
†Thalassemydidae	†Neusticemys †Palaeomedusa †Thalassemys
	†Adocus †Angolachelys †Argillochelys †Bashuchelys †Brodiechelys †Ctenochelys †Compsemys †Hoyasemys †Larachelus †Leyvachelys †Plesiochelys †Toxochelys †Uluops †Xinjiangchelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Cearachelys †Chedighaii †Chupacabrachelys †Foxemys †Galianemys †Kinkonychelys †Kurmademys †Polysternon †Puentemys †Sankuchemys
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Latentemys Peltocephalus Podocnemis †Stupendemys

	†Caribemys †Caririemys †Chubutemys †Hispaniachelys †Tacuarembemys

Phylogenetic arrangement of turtles based on turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status. † = extinct.

See also List of Testudines families

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Taxon identifiers
Gamerabaena	Wikidata: Q20677867 EoL: 18757363 Fossilworks: 202998 IRMNG: 1473043
Gamerabaena sonsalla	Wikidata: Q5520054 Fossilworks: 202999