Bombardier Electrostar

357030 at Barking in National Express c2c livery. Electrostar trains are the new standard on many of London's commuter routes.

c2c uses Class 357 and Class 387 units on services down the London, Tilbury and Southend Railway line from Shoeburyness and Southend to London Fenchurch Street.

A Southeastern Class 375 Electrostar at Strood.

Interior of a Southeastern electrostar (Class 375).

The Class 375 is the backbone of Southeastern's long-distance routes, seeing services on most of its lines originating from its London termini (London Victoria, Charing Cross, Cannon Street and London Bridge) including;

• Chatham Main Line
  • Maidstone East Line
• Medway Valley Line
• South Eastern Main Line
  • Hastings Line
  • Ashford to Ramsgate (via Canterbury West) line
  • Kent Coast Line
• North Kent Line
• Mid Kent Line

On the outer suburban portions of these above routes, Class 465/9 Networkers support the Electrostars, but they do not work in multiple together.

A Southeastern Class 376 Electrostar in Southeastern livery.

The Class 376 operates on the metro routes in suburban London, in conjunction with the Class 465 and Class 466 Networkers, operating over the London portion of the above lines from the London Termini (including Blackfriars) out to Dartford and Sevenoaks);

• North Kent Line (to Dartford)
• Bexleyheath Line (to Dartford)
• Dartford Loop Line (to Dartford)
• Chatham Main Line (to Sevenoaks)
• Maidstone East Line (to Sevenoaks)
• South Eastern Main Line (to Sevenoaks)
• Hayes Line

This leaves the Bromley North Line and Sheerness Line, both operated by Class 466s (2 car Networkers) which also used to operate on the Medway Valley Line prior to the May 2012 Timetable Changes.

A Southern Class 377.

Southern's Class 377 fleet is found on all parts of the network apart from the non-electrified routes. They also now run frequently in metro routes alongside the Class 455s and used to run alongside the Class 456s until their transfer to South West Trains in 2014.

• Brighton Main Line (Victoria–Gatwick-Brighton)
• East Coastway (Brighton–Eastbourne/Hastings)
• West Coastway (Brighton–Portsmouth/Southampton)
• Arun Valley Line (Victoria–Horsham/Littlehampton/Chichester)
• West London Line (East Croydon–Milton Keynes Central) (Using Class 377/2 or Dual Voltage Class 377/7)

• London Victoria–Horsham via Dorking
• London Victoria–East Grinstead
• London Bridge–Horsham via East Croydon
• London Victoria-Reigate
• Redhill-Tonbridge

Often found on

• London Victoria–Dorking via Sutton
• London Victoria–Epsom Downs
• London Bridge-London Victoria via Sydenham
• London Bridge-Caterham
• London Victoria-Caterham
• London Victoria-Epsom
• London Bridge-Tattenham Corner

A London Overground Class 378 in TfL livery at Crystal Palace.

London Overground operates Class 378s over five lines around London:

• North London Line (Richmond–Stratford)
• West London Line (Clapham Junction-Willesden Junction/Stratford via West Brompton)
• Watford DC Line (Euston–Watford Junction)
• East London Line (Crystal Palace/New Cross/West Croydon–Dalston Junction/Highbury & Islington via Canada Water)
• South London Line (Clapham Junction-Dalston Junction)

The sixth major route it's responsible for is the unelectrified Gospel Oak to Barking Line. For this, London Overground obtained Class 172 Turbostar DMUs and have been replaced by new Class 710 Aventra EMUs.

On 8 June 2010, the route between Sandton and OR Tambo International Airport in South Africa opened in time for the 2010 FIFA World Cup.[6] The rest between Johannesburg Park Station and Rosebank was to be completed in 2011. This section was actually opened 7 June 2012,[7] the delay caused by work to resolve a water-seepage problem in the single-track tunnel section between Rosebank and Park. Although railways in South Africa use the 1,067 mm (3 ft 6 in) Cape gauge, Gautrain is built to the more expensive standard gauge of 1,435 mm (4 ft 8 ¹⁄₂ in). According to the Gautrain planning and implementation study,[8] this is done for several reasons, including that standard gauge is safer and more comfortable to passengers. The rolling stock is also easier, quicker and less expensive to obtain than Cape Gauge rolling stock, and standard gauge is also less expensive to maintain as it is more tolerant of track imperfections than Cape Gauge. Standard gauge allows for travel at Gautrain's required speed of 160 km/h (99 mph).

Abellio Greater Anglia Class 379 "Electrostar" EMU 379014 arrives at Tottenham Hale with a service to Liverpool Street.

From March 2011 National Express East Anglia introduced 30 x 4 car Class 379s on Stansted Express and West Anglia Main Line services.[9] These incorporate some features of Bombardier's planned 'Aventra' Mark 2 Electrostars.[5]

All units had entered service by mid-August 2011, two months ahead of schedule. A major timetable update in December 2011 entailed the introduction of 12-car trains on some peak workings to and from Cambridge.[10] These trains are now operated by Greater Anglia as of 5 February 2012.

All of Greater Anglia's units will be replaced in 2020 by Class 745s on the Stansted Express route and Class 720s on the Cambridge route.

Great Western Railway Class 387 "Electrostar" EMUs 387131 & 387132 at London Paddington on 2 September 2016.

From September 2016 Great Western Railway introduced 45 4 car Class 387/1s on peak services between London Paddington and Hayes & Harlington. They currently replace the Class 165s and the Class 166s on the Thames Valley services and now operate between London Paddington and Reading, Didcot Parkway and Newbury.

GWR announced in 2018 that they will modify 12 units to be used on Heathrow Express due to the Class 332s no longer having a depot.

From December 2019, services to Reading have mainly transferred to TfL Rail as part of the Elizabeth Line.

From late 2016, 29 of the Class 387/1s operating on Thameslink were displaced by the delivery of Class 700 Desiro City units, and were transferred to Great Northern. They operate mostly on the Kings Cross-Cambridge-King's Lynn route, though they can also been seen on other services. These units were delivered in the livery of Southern, with green doors and Southern upholstery.