Cyrtodactylus

Cyrtodactylus (Greek κυρτος kurtos "curved", from κυπτω kuptō "to stoop"; δακτυλος daktulos "finger, toe") is a diverse genus of Asian geckos, commonly known as bent-toed geckos or bow-fingered geckos. It has at least 250 described species at present,[1] which makes it the largest of all gecko genera.

Cyrtodactylus
Marbled bow-fingered gecko (Cyrtodactylus marmoratus)
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Squamata
Family: Gekkonidae
Subfamily: Gekkoninae
Genus: Cyrtodactylus
Gray, 1827
Species

At least 250 recognized species, see article.

Description

Instead of possessing dilated digits like other geckos, members of Cyrtodactylus have slender, curved toes to which the common names for the genus can be attributed.

Taxonomy

The phylogenetic relationships with this genus and the genus Geckoella has not been resolved to date. Based on morphology, Bauer (2002) suggested that it was a subgenus of Cyrtodactylus but a phylogenetic study by Feng et al. (2007), though with inadequate sampling of taxa, restored it to generic status once again.[2] All species of Geckoella have been placed here pending a more definitive study.

Species

More than 250 species described. In 2019, six new species of the Cyrtodactylus intermedius complex were identified in the Cardamom Mountains and Associated Highlands region of Southeast Asia.[3]

Species Authority Common name Geographic range
C. aaroniGünther & Rösler, 2003[4]New Guinea
C. adleriDas, 1997Nicobar Islands, Andaman Sea, India
C. adorusShea, Couper, Wilmer& Amey, 2011 [5]Queensland, Australia
C. aequalisBauer, 2003[6]Mon State, Myanmar
C. agamensis(Bleeker, 1860)Agam bow-fingered geckoSumatra, Indonesia
C. agusanensisTaylor, 1915Mindanao bow-fingered geckoMindanao Island, The Philippines
C. albofasciatus(Boulenger, 1885)Boulenger's Indian geckowestern Ghats, India
C. angularis(M.A. Smith, 1921)angulated bow-fingered geckoEastern Thailand
C. annandaleiBauer, 2003[6]Annandale's leaf-toed geckoSagaing Division, Myanmar
C. annulatusTaylor, 1915[7]Annulated bow-fingered geckoMindanao Island, The Philippines
C. arcanusOliver, Richards & Sistrom 2012 [8][9]Papua New Guinea
C. astrumGrismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels, 2012 [10][11]Starry bent-toed geckoSouthern Thailand, Peninsular Malaysia
C. atremus Kraus & Weijola, 2019Papua New Guinea
C. aungliniMurdoch, Grismer, Wood, Thura, Win, Grismer, Trueblood, & Quah, 2018[3]Kyauk Nagar Cave bent-toed geckoMyanmar
C. auralensisMurdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[3]Phnom Aural bent-toed geckoPhnom Aural, Cambodia
C. aurensisGrismer, 2005[12]Aur Island bent-toed geckoAur Island, Malaysia
C. auribalteatusSumontha, Panitvong & Deein, 2010[13]Golden-belted bent-toed geckoPhitsanulok, Thailand
C. australotitiwangsaensisGrismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels, 2012 [14]Southern Titiwangsa bent-toed geckoPeninsular Malaysia
C. ayeyarwadyensisBauer, 2003[6]Rakhine State, Myanmar
C. badenensisNguyen, Orlov & Darevsky, 2006[15]Southern Vietnam
C. baluensisMocquard, 1890[16]Balu bow-fingered geckoMount Kinabalu, Borneo
C. bansocensisLuu, Nguyen, Le, Bonkowski & Ziegler, 2016[17]Ban Soc bent-toed geckoLaos
C. batikIskandar, Rachmansah & Umilaela, 2011[18]Batik bent-toed geckoIndonesia
C. battalensisKhan, 1993Reticulate plump-bodied geckoPakistan
C. batucolusL.L. Grismer, Onn, J.L. Grismer, Wood & Belabut, 2008[19]Besar Island bent-toed geckoPulau Besar of the Water Islands Archipelago off the west coast of Melaka, Peninsular Malaysia
C. bayinnyiensisGrismer, Wood Jr., Thura, Quah, Murdoch, Grismer, Herr, Lin, & Kyaw, 2018Bayin Nyi Cave bent-toed geckoMyanmar
C. bhupathyiAgarwal, Mahony, Giri, Chaitanya, & Bauer, 2018[20]Northeast India
C. bichnganaeTri & Grismer, 2010Bich Ngan's bent-toed geckoVietnam
C. bidoupimontisNazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler, 2012[21]Vietnam
C. bintangrendahGrismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels, 2012[14]Bintang lowland bent-toed geckoPeninsular Malaysia
C. bintangtinggiGrismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels, 2012[14]Peninsular Malaysia
C. biordinisBrown & Mccoy, 1980[22]Guadalcanal bow-fingered geckoMount Austen, Guadalcanal, Solomon Islands
C. bobroviNguyen et al., 2015[23]Bobrov's bent-toed geckoVietnam
C. bokorensisMurdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[3]Bokor Plateau, Cambodia
C. boreoclivusOliver, Krey, Mumpuni & Richards, 2011[24]Papua New Guinea
C. brevidactylusBauer, 2002[25]Myanmar
C. brevipalmatus(M.A. Smith, 1923)[26]Kampuchea bow-fingered gecko, short-hand forest geckoKhao Luang, Nakhon Si Thammarat, Peninsular Thailand
C. buchardiDavid, Teynie & Ohler, 2004[27]Laos
C. bugiamapensisNazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler, 2012[21]Vietnam
C. calameiLuu, Bonkowki, Nguyen, Le, Schneider, Ngo, & Ziegler, 2016Calame's bent-toed geckocentral Laos
C. caovansungi Orlov & Quang Truong, 2007 [28]Vietnam
C. capreoloidesRösler, Richards & Günther, 2007Papua New Guinea
C. cardamomensisMurdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[3]Cardamom Mountains bent-toed geckoCardamom Mountains, Cambodia
C. cattienensisGeissler, Nazarov, Orlov, Böhme, Phung, Nguyen & Ziegler, 2009[29]Cattien bent-toed geckoVietnam
C. cavernicolusInger & King, 1962Sarawak bow-fingered gecko, Niah Cave geckoSarawak
C. cayuensis Li, 2007Cayu bent-toed geckoChina (Cayu County, Xizang)
C. celatusKathriner et al., 2014[30]Indonesia
C. chambaAgarwaL, Khandekar, & Bauer, 2018Chamba bent-toed geckoHimachal Pradesh, India
C. chanhomeaeBauer, Sumontha & Pauwels, 2003[31]Thailand
C. chaunghanakwaensisGrismer, Wood Jr., Thura, Quah, Murdoch, Grismer, Herr, Lin, & Kyaw, 2018Chaunghanakwa Hill bent-toed geckoMyanmar
C. chauquangensisQuang, Orlov, et al. 2007[32]Vietnam
C. chrysopylosBauer, 2003[6]Panluang-Pyadakin Wildlife Sanctuary, Shan State, Myanmar
C. collegalensis(Beddome, 1870)forest spotted gecko, Kollegal ground geckoIndia
C. condorensis(M.A. Smith, 1921)Pulo Condore bow-fingered gecko, Con Dao bow-fingered geckoPulo Condore Island, Vietnam
C. consobrinoidesAnnandale, 1905southern Myanmar
C. consobrinusW. Peters, 1871Peters's bow-fingered gecko, thin-banded forest geckoMalaysian peninsula, Borneo
C. cracensBatuwita & Bahir, 2005Sri Lanka
C.. crustulusOliver, Hartman, Turner, Wilde, Austin, & Richards, 2020Manus bent-toed geckoPapua New Guinea (Manus Island)
C. cryptusHeidrich, Rösler, Thanh, Böhme & Ziegler, 2007[33]Vietnam
C. cucdongensisSchneider & Phung TM, 2014[34]Cucdong bent-toed geckoVietnam
C. cucphuongensis Ngo & Onn, 2011[35]Vietnam
C. dammathetensisGrismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[36]Dammathet cave bent-toed geckoMyanmar
C. darevskiiNazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov, & Chulisov, 2014 Darevski's bent-toed geckoLaos
C. darmandvillei(Weber, 1890) Darmandville bow-fingered geckoFlores, Komodo, Pulau Kalao, Sumbawa, Sikka, Lombok in Indonesia
C. datiTri, 2013[37]Dat bent-toed geckoVietnam
C. dattanensis(Khan, 1980)plump banded gecko, Khan's bow-fingered geckonw Pakistan
C. dattkyaikensisGrismer, Wood, Quah, Grismer, Thura, Oaks, & Lin, 2020Datt Kyaik Hill bent-toed geckoMyanmar
C. dayangbuntingensisQuah, Grismer, Wood Jr., & Sah, 2019peninsular Malaysia
C. deccanensis(Günther, 1864)Günther's Indian geckoNorthern Western Ghats of India
C. derongoBrown & Parker, 1973Derongo bow-fingered geckoPapua, New Guinea
C. deveti(Brongersma, 1948)Moluccan bow-fingered geckoMoluccas, Indonesia
C. doisuthepKunya, K. et al. 2014[38]Doi Suthep bent-toed geckoThailand
C. dumnuiiBauer, Kunya, Sumontha, Niyomwan, Pauwels, Chanhome & Kunya, 2010[39]Dumnui's bent-toed geckoThailand
C. durioGrismer, Anuar, Quah, Muin, Onn, Grismer & Ahmad, 2010[40]Durian bent-toed geckoMalaysia
C. edwardtayloriBatuwita & Bahir, 2005Sri Lanka
C. eisenmanaeTri, 2008[41]Eisenman's bent-toed geckoVietnam
C. elokDring, 1979Malaysia bow-fingered geckowestern Malaysia
C. epiroticusKraus, 2008Papua New Guinea
C. equestrisOliver, Richards, Mumpuni & Rosler, 2016 Papua New Guinea
C. erythropsBauer, Kunya, Sumontha, Niyomwan, Panitvong, Pauwels, Chanhome & Kunya, 2009[42]Red-eyed bent-toed geckoThailand
C. evanquahiWood, Grismer, Muin, Anuar, Oaks, & Sites, 2020Evan Quah’s banded bent-toed geckopeninsular Malaysia
C. fasciolatus(Blyth, 1861)[43]banded bent-toed geckoNorthwest India
C. feae(Boulenger, 1893)Myanmar
C. fraenatus(Günther, 1864)Sri Lanka bow-fingered geckoSri Lanka
C. fumosus(Müller, 1895)[44]Sulawesi, Indonesia
C. gansiBauer, 2003[6]Min Dat District, Chin State, Myanmar
C. gialaiensisLuu, Dung, Nguyen, Le, & Ziegler, 2017 Vietnam
C. gordongekkoiDas, 1994 [45]Lombok, Indonesia
C. grismeriTri, 2008Grismer's bent-toed geckoVietnam
C. guakanthanensisGrismer LL, Belabut DM, 2014[46]Gua Kanthan bent-toed geckoMalaysia
C. gubaotWelton, Siler, Linkem, Diesmos & Brown, 2010[47]the Philippines
C. gubernatoris(Annandale, 1913)Sikkimese bent-toed geckoDarjeeling, India
C. gunungsenyumensisGrismer et al., 2016[48]Gunung Senyum bent-toed geckoMalaysia
C. guwahatiensisAgarwal, Mahony, Giri, Chaitanya, & Bauer, 2018India
C. halmahericusMertens, 1929 [49]Indonesia
C. hidupselamanyaGrismer, Wood, Anuar, Grismer, Quah, Murdoch, Muin, Davis, Auilar, Klabacka, Cobos, Aowphol, & Sites, 2016 Chiku bent-toed geckopeninsular Malaysia
C. hikidaiRiyanto A, 2012[50]Bunguran island, Indonesia
C. himalayanusDuda & Sani, 1977Himalaya bow-fingered geckoJammu, India
C. himalayicus(Annandale, 1906)Himalaya bent-toed geckoIndia (West Bengal, Darjeeling)
C. hinnamnoensisLuu, Bonkowski, Nguyen, Le, Schneider, Ngo, & Ziegler, 2016Hinnamno bent-toed geckoLaos
C. hitchi, Riyanto, Kurniati & Engilis, 2016southeastern Sulawesi, Indonesia
C. hontreensisTri, Grismer & Grismer, 2008[51]Hon Tre bent-toed geckoVietnam
C. hoskiniShea, Couper, Wilmer & Amey, 2011[52]Queensland, Australia
C. huongsonensisLuu, Do & Ziegler, 2011[53]Vietnam
C. huynhiNgo & Bauer, 2008[54]Huynh's bent-toed geckoVietnam
C. ingeriHikida, 1990Sabah bow-fingered geckoSabah, Borneo
C. interdigitalis(Ulber, 1992)skin-toe forest geckoThailand, Laos
C. intermediusUlber, 1993intermediate bow-fingered geckoCambodia, Vietnam, and Thailand
C. inthanonKunya et al., 2015[55]Doi Inthanon bent-toed geckoThailand
C. irianjayaensisRossler, 2001Papua, New Guinea
C. irregularis(M.A. Smith, 1921)irregular bow-fingered geckoVietnam
C. jaegeriLuu VQ, Calame T, 2014[56]Laos
C. jaintiaensisAgarwal, Mahony, Giri, Chaitanya, & Bauer, 2018India
C. jambanganWelton, Siler, Diesmos & Brown, 2010[57]the Philippines
C. jarakensisL.L. Grismer, Onn, J.L. Grismer, Wood & Belabut, 2008[19]Jarak Island bent-toed geckoPulau Jarak off the west coast of Perak, Peninsular Malaysia
C. jarujini(Ulber, 1992)Jarujin's forest geckoLaos and northern Thailand
C. jelawangensisAnuar S, Quah ES, 2014[58]Jelawang bent-toed geckoThai-Malay Peninsula
C. jellesmaeBoulenger, 1897Kabaena bow-fingered geckoMasarang in Indonesia
C. jeyporensis(Beddome, 1877)Jeypore rock geckoeastern Ghats, India
C. kazirangaensisAgarwal, Mahony, Giri, Chaitanya, & Bauer, 2018India
C. khammouanensisNazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov, & Chulisov, 2014Khammouane bent-toed geckoLaos
C. khasiensis(Jerdon, 1870)Khasi Hills bent-toed geckoIndia and northern Burma
C. khelangensisPauwels OS, Sumontha M, 2014[59]Lampang bent-toed geckoThailand
C. kimberleyensisAaron & Doughty, 2012[60]Kimberley bent-toed gecko, East Montalivet Island geckoAustralia
C. kingsadaiZiegler, Phung, Le & Nguyen, 2013[61]Vietnam
C. klakahensisHartmann, Mecke, Kieckbusch, Mader, & Kaiser, 2016Klakah bent-toed geckoJava, Indonesia
C. klugeiKraus, 2008Papua New Guinea
C. kunyaiPauwels, Sumontha, Keeratikiat & Phanamphon, 2014[62]Kunya's bent-toed geckoThailand
C. laangensisMurdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[3]Phnom Laang bent-toed geckoPhnom Laang, Cambodia
C. laevigatusDarevsky, 1964Komodo Island
C. langkawiensisGrismer, Wood, 2012[14]Langkawi Island bent-toed geckoLangkawi, Malaysia
C. lateralisWerner, 1896Sumatra bow-fingered geckoSumatra
C. lawderanus(Stoliczka, 1871)Lawder's bent-toed geckonorthwestern India
C. leegrismeriChan & Norhayati, 2010[63]Tenggol Island bent-toed geckoMalaysia
C. lekaguliGrismer et al., 2012[14]Southern Thailand
C. lenggongensisGrismer, Wood, Anuar, Grismer, Quah, Murdoch, Muin, Davis, Auilar, Klabacka, Cobos, Aowphol, & Sites, 2016Lenggong bent-toed geckopeninsular Malaysia
C. lenyaMulcahy, Thura, & Zug, 2017Lenya banded bent-toed geckoMyanmar
C. limajalurDavis, Bauer, Jackman, Nashriq, & Das, 2019five-banded bent-toed geckoEast Malaysia
C. linnoensisGrismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[36]Linno Cave bent-toed geckoMyanmar
C. linnwayensisGrismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[36]Linn-Way bent-toed geckoMyanmar
C. lomyenensisNgo & Pauwels, 2010[64]Lomyen bent-toed geckoLaos
C. loriae(Boulenger, 1897)Boulenger's bow-fingered geckoPapua New Guinea
C. louisiadensis(DeVis, 1892)ring-tailed geckoLouisiade Archipelago, Papua New Guinea
C. macrotuberculatusGrismer & Ahmad, 2008[65]tuberculate bent-toed gecko, large tubercled bent-toed geckoMalaysia
C. majulahGrismer, Wood & Lim 2012Singapore; Indonesia
C. malayanusDe Rooij, 1915Borneo bow-fingered geckoBorneo
C. mamanwaWelton, Siler, Linkem, Diesmons & Brown 2010[47][66]The Philippines
C. mandalayensisMahony 2009[67]Myanmar
C. manosOliver, Karkkainen, Rosler, & Richards, 2019yellow-snouted bent-toed geckoPapua New Guinea
C. markuscombaiiDarevsky, Helfenberger, Orlov & Shah, 1998[68]striped geckoNepal
C. marmoratusGray, 1831marbled bow-fingered geckoThailand, Malay Peninsula, Indonesia, Papua New Guinea.
C. martiniTri, 2011[69]Martin's bent-toed geckoVietnam
C. martinstolliDarevsky, Helfenberger, Orlov & Shah, 1998[68]barred geckoNepal
C. matsuiiHikida, 1990Hikida's bow-fingered gecko,
Matsui's bent-toed gecko		Sabah, Borneo
C. mcdonaldiShea, Couper, Wilwer & Amey 2011 [70]Australia
C. medioclivusOliver, Richards & Sistrom, 2012 [71]Papua New Guinea
C. meersiGrismer, Wood, Quah, Murdoch, M. Grismer, Herr, Espinoza, Brown, & Lin, 2018Bago Yoma bent-toed geckoMyanmar (Yangon)
C. metropolisGrismer LL, Wood PJ, Onn CK, Anuar S, Muin MA, 2014[72]Malaysia
C. mimikanus(Boulenger, 1914)false bow-fingered gecko, Mimika bow-fingered gecko, Mimika bent-toed geckoPapua New Guinea
C. minorOliver & Richards, 2012[73]Papua New Guinea
C. mombergiGrismer, Wood, Quah, Thura, Herr, & Lin, 2019Indawgyi bent-toed geckoMyanmar
C. montanusAgarwal, Mahony, Giri, Chaitanya, & Bauer, 2018India
C. muangfuangensisSitthivong, Luu, Ha, Nguyen, Le, & Ziegler, 2019Muangfuang bent-toed geckoLaos
C. multiporusNazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov, & Chulisov, 2014Multipored bent-toed geckoLaos
C. muluensisDavis, Bauer, Jackman, Nashriq, & Das, 2019Mulu bent-toed geckoEast Malaysia
C. muruaKraus & Allison, 2006[74]Papua New Guinea
C. myaleiktaungGrismer, Wood, Thura, Win, Grismer, Trueblood, & Quah, 2018Mya Leik Taung bent-toed geckoMyanmar
C. myintkyawthuraiGrismer, Wood, Quah, Murdoch, M. Grismer, Herr, Espinoza, Brown, & Lin, 2018Mt. Popa bent-toed geckoMyanmar (Mandalay region, Bago Yoma Range, Bago Region)
C. nagalandensisAgarwal, Mahony, Giri, Chaitanya, & Bauer, 2018India
C. naungkayaingensisGrismer, Wood Jr., Thura, Quah, Murdoch, Grismer, Herr, Lin, & Kyaw, 2018Myanmar
C. nebulosus(Beddome, 1870)Clouded Indian geckoIndia
C. nepalensisSchleich & Kästle, 1998[75]Nepalese rock geckoNepal
C. nigriocularisNguyễn, Orlov & Darevsky, 2007southern Vietnam
C. novaeguineae(Schlegel, 1844)New Guinea bow-fingered geckoNew Guinea
C. nuauluOliver, Edgar, Mumpuni, Iskandar & Lilley, 2009[76]Indonesia
C. nyinyikyawiGrismer, Wood, Thura, Win, & Quah, 2019Shwe Settaw bent-toed geckoMyanmar
C. oldhami(Theobald, 1876)Oldham's bow-fingered geckoMyanmar and Thailand
C. otaiNguyen et al., 2015[23]Ota's bent-toed geckoVietnam
C. pageliSchneider et al., 2011[77]Laos
C. pantiensisL.L. Grismer, Onn, J.L. Grismer, Wood & Belabut, 2008[19]Panti Mountain bent-toed geckoGunung Panti Forest Reserve, Johor, Peninsular Malaysia
C. papilionoidesUlber & Grossmann, 1991butterfly forest geckoThailand
C. papuensis(Brongersma, 1934)Papua bow-fingered geckoPapua New Guinea
C. paradoxusDarevsky & Szczerbak, 1997Kieng Giang, Phu Quoc, and Hon Thom Islands in Vietnam
C. payacolaJohnson, Quah Anuar, Muin, Wood, Grismer, Greer, Onn, Ahmad, Bauer& Grismer 2012 [78]Malaysia
C. pyadalinensisGrismer, Wood, Thura, Win, & Quah, 2019Myanmar
C. payarhtanensisMulcahy, Thura, & Zug, 2017Tenasserim Mountain bent-toed geckosouthern Myanmar
C. peguensis(Boulenger, 1893)Thai bow-fingered gecko, Pegu forest geckoThailand, western Malaysia, and Myanmar
C. petaniRiyanto, Grismer & Wood, 2015[79]Farmer's bent-toed geckoIndonesia
C. pharbaungensisGrismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[36]Pharbaung Cave bent-toed geckoMyanmar
C. phetchaburiensisPawels, Sumontha, & Bauer, 2016Thailand
C. philippinicusSteindachner, 1867Philippine bow-fingered geckonorthern Philippines
C. phnomchiensisNyang, Henson & Stuart, 2020[80]Prey Lang bent-toed gecko[81]Prey Lang, Cambodia
C. phongnhakebangensisZiegler, Rösler, Herrmann, and Vũ, 2002Phong Nha-Ke Bang bow-fingered geckoVietnam
C. phuketensisSumontha, Pawels, Kunya, Nitikul, Samphanthamit, & Grismer, 2012 [82]Phuket bent-toed geckoPhuket, Thailand
C. phuocbinhensisNguyen, Le, Tran, Orlov, 2013 [83]Vietnam
C. phuquocensisTri, Grismer & Grismer, 2010[84]Phu Quoc bent–toed geckoVietnam
C. pinlaungensisGrismer, Wood, Quah, Thura, Oaks, & Lin, 2019Pinlaung bent-toed geckoMyanmar
C. pronarusShea, Couper, Wilmer & Amey, 2011[52]Australia
C. psaropsHarvey et al., 2015[85]Speckle-faced bent-toed geckoIndonesia
C. pseudoquadrivirgatus(Rösler, Nguyễn, Vũ, Ngô & Ziegler, 2008)central Vietnam
C. pubisulcus Inger, 1958Inger's bow-fingered geckoBorneo
C. puhuensisNguyen, Yang, Thi Le, Nguyen, Orlov, Hoang, Nguyen, Jin, Rao, Hoang, Che, Murphy & Zhang, 2014[86]Pù Hu bent-toed geckoVietnam
C. pulchellusGray, 1827Malayan forest geckoPenang, Malaysia
C. pyinyaungensisGrismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[36]Pyinyaung bent-toed geckoMyanmar
C. quadrivirgatusTaylor, 1962Taylor's bow-fingered gecko, four-striped forest gecko, marbled bent-toed geckoThailand, Malaysia, Singapore, and Indonesia
C. rambodaBatuwita & Bahir, 2005Sri Lanka
C. ranongensisSumontha, Pauwels, Panitvong, Kunya & Grismer, 2015[87]Ranong bent-toed geckoThailand
C. redimiculusKing, 1962Palawan bow-fingered geckoPalawan, Philippines
C. rexOliver, Richards, Mumpuni, & Rosler, 2016Papua New Guinea
C. rishivalleyensisAgarwal, 2016Rishi Valley geckoellaIndia
C. robustusKraus, 2008Papua New Guinea
C. roesleriZiegler, Nazarov, Orlov, Nguyen, Vu, Dang, Dinh, & Schmitz, 2010 [88]Roesler's bent-toed geckoVietnam & Laos
C. rosichonariefiRiyanto, Grismer & Wood, 2015[89]Rosichonariefi's bent-toed geckoIndonesia
C. rubidus(Blyth, 1861)red bow-fingered geckoAndaman Islands; Myanmar
C. ruffordLuu et al., 2016[90]Rufford bent-toed geckoLaos
C. russelliBauer, 2003[6]Htamanthi Wildlife Sanctuary, Sagaing Division and Indawgyi Wildlife Sanctuary, Rakhine State, Myanmar
C. sadanensisGrismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[36]Sadan Cave bent-toed geckoMyanmar
C. sadansinensisGrismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[36]Sadan Sin Cave bent-toed geckoMyanmar
C. sadleiriWells & Wellington, 1984Sadleir's bow-fingered geckoChristmas Island
C. saiyokPanitvong, Sumontha, Tunprasert & Pauwels, 2014[91]Sai Yok bent-toed geckoThailand
C. salomonensisRösler, Richards & Günther, 2007[92]Solomon Islands
C. samroiyotPauwels OS, Sumontha M, 2014[93]Sam Roi Yot bent-toed geckoThailand
C. sangiPauwels, Nazarov, Bobrov, & Poyarkov, 2018southern Vietnam
C. sanookPauwels, Sumontha, Latinne & Grismer, 2013[94]Sanook bent-toed geckoThailand
C. sanpelensisGrismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[36]Sanpel Cave bent-toed geckoMyanmar
C. semenanjungensisGrismer & Leong, 2005Peninsular bent-toed geckowestern Malaysia
C. semiadiiRiyanto A, Bauer AM, Yudha DS, 2014[95]Indonesia
C. semicinctusHarvey et al., 2015[85]Kerinci bent-toed geckoIndonesia
C. septentrionalisAgarwal, Mahony, Giri, Chaitanya, & Bauer, 2018India
C. septimontiumMurdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[3]Bảy Núi bent-toed geckoBảy Núi Hills, Vietnam
C. seribuatensisYoumans & Grismer, 2006Seribuat bent-toed geckowestern Malaysia
C. sermowaiensis(de Rooij, 1915)DeRooij's bow-fingered geckoNew Guinea
C. serratusKraus, 2007[96]Papua New Guinea
C. sharkariGrismer LL, jr. et al., 2014[58]Sharkari's bent-toed geckoThai-Malay Peninsula
C. shwetaungorumGrismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[36]Shwetaung bent-toed geckoMyanmar
C. sinyineensisGrismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[36]Sin Yine Cave bent-toed geckoMyanmar
C. slowinskiiBauer, 2002Myanmar
C. sobaBatuwita & Bahir, 2005Dumbara bent-toed gecko, Knuckles bent-toed geckoSri Lanka
C. sommerladiLuu, Bonkowski, Nguyen, Le, Schneider, Ngo, & Ziegler, 2016Sommerlad's bent-toed geckocentral Laos
C. soniLe, Nguyen, Le, & Ziegler, 2016Son's bent-toed geckoVietnam
C. sonlaensisNguyen, Pham, Ziegler, Ngo, & Le, 2017northwestern Vietnam
C. soudthichakiLuu, Calame, Nguyen, Bonkowski & Ziegler, 2015[97]Soudthichak's bent-toed geckoLaos
C. speciosus(Beddome, 1870)forest spotted geckoIndia
C. spelaeusNazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov, & Chulisov, 2014Laos
C. spinosusLinkem, McGuire, Hayden, Setiadi, Bickford & Brown, 2008Sulawesi, Indonesia
C. srilekhaeAgarwal, 2016Bangalore geckoellaIndia
C. stresemanni Rösler & Glaw, 2008Stresemann's bent-toed geckoMalaysia
C. subsolanusBatuwita & Bahir, 2005Sinharaja bent-toed geckoSri Lanka
C. sumonthaiBauer, 2002Sumontha's geckoThailand
C. sumuroiWelton, Siler, Linkem, Diesmos & Brown, 2010[47]the Philippines
C. surinChan-ard & Makchai (2011)[98]Thailand
C. sworderi(M.A. Smith, 1925)Johore bow-fingered gecko, Kota-tinggi forest gecko, Sworder's bent-toed geckoMalaysia
C. tahunaRiyanto, Arida, & Koch, 2018[99]Sangir Island, Indonesia
C. takouensisVan Tri N, Gamble T., 2010[100]Takou bent-toed geckoVietnam
C. tamaiensis(M.A. Smith, 1940)[101]Tamai bent-toed geckoMyanmar
C. tamboraRiyanto, Mulyadi, McGuire, Kusrini, Febylasmia, Basyir, & Kiser, 2017Tambora bent-toed geckoSumbawa, Indonesia
C. tanahjampeaRiyanto, Hamidy, & McGuire, 2018Tanahjampea bent-toed geckoTanahjampea Island, Indonesia
C. tanimNielsen & Oliver, 2017Papua New Guinea
C. taungwineensisGrismer, Wood, Quah, Grismer, Thura, Oaks, & Lin, 2020Taung Wine Hill bent-toed geckoMyanmar
C. tautbatorumWelton, Siler, Diesmos & Brown, 2009[102]the Philippines
C. taybacensisPham, Le, Ngo, Ziegler, & Nguyen, 2019Taybac bent-toed geckoVietnam
C. taynguyenensisNguyen, Le, Tran, Orlov, Lathrop, Macculloch, Le, Jin, Nguyen, Nguyen, Hoang, Che, Murphy & Zhang 2013[103]Vietnam
C. tebuensisGrismer, Anuar, Muin, Quah & Wood, 2013[104]Tebu Mountain bent-toed geckoMalaysia
C. teynieiDavid, Nguyen, Schneider & Ziegler, 2011[105]Laos
C. thathomensisNazarov, Pauwels, Konstantinov, Chulisov, Orlov, & Poyarkov, 2018Thathom bent-toed geckoLaos
C. thirakhuptiPauwels, Bauer, Sumontha & Chanhome, 2004[106]Thirakhupt's bent-toed geckoThailand
C. thochuensisNgo, Van Tri; Grismer, L. Lee 2012[107]Thochu bent–toed geckoTho Chu Island, Vietnam
C. thuongaePhung TM & Schingen MV, 2014[108]Vietnam
C. thylacodactylusMurdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[3]Phnom Dalai, Cambodia
C. tibetanusBoulenger, 1905Tibet
C. tigroidesBauer, Sumontha & Pauwels, 2003[31]Tiger bent-toed geckoThailand
C. timurMuin MA, Mohamed M, Onn CK, Sumarli AX, Loredo AI, Heinz HM 2014[58]Banjaran Timur bent-toed geckoThai-Malay Peninsula
C. tiomanensisDas & Lim, 2000[109]Tioman Island bent-toed geckoMalaysia
C. triedrus(Günther, 1864)spotted bent-toed gecko, Sri Lanka gecko, spotted bow-fingered gecko, spotted ground geckoSri Lanka
C. trilatofasciatusGrismer & Wood Jr., 2012[110]Cameron Highlands bent-toed geckoMalaysia
C. tripartitusKraus, 2008Papua New Guinea
C. tripuraensisAgarwal, Mahony, Giri, Chaitanya, & Bauer, 2018India
C. tuberculatusLucas & Frost, 1900Queensland, Australia
C. urbanus Purkayastha, Das, Bohra, Bauer, & Agarwal, 2020urban bent-toed geckoIndia (Assam)
C. varadgiriiAgarwal et al., 2016[111]Giri's GeckoellaMumbai, India
C. variegatusBlyth, 1859variegated bow-fingered gecko, Moulmein forest geckoMyanmar and Thailand
C. vilaphongiSchneider, Nguyen, Le, Nophaseud, Bonkowski & Ziegler, 2014Laos
C. wakeorumBauer, 2003[6]Southern Rakhine State, Myanmar
C. wallaceiHayden, Brown, Gillespie, Setiadi, Linkem, Iskandar, Umilaela, Bickford, Riyanto, Mumpuni & McGuire,2008Sulawesi, Indonesia
C. wangkulangkulaeSumontha, M. et al. 2014[112]Wangkulangkul's bent-toed geckoThailand
C. wayakonei Nguyen, Kingsada, Rösler, Auer & Ziegler, 2010[113]Northern Laos & Yunnan, China [114]
C. welpyanensisGrismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[36]Wel Pyan Cave bent-toed geckoMyanmar
C. wetariensisDunn, 1927Wetar bow-fingered geckoWetar, Indonesia
C. yakhuna(Deraniyagala, 1945)northern Sri Lanka gecko, spotted bow-fingered gecko, blotch bow-fingered gecko, demon geckoSri Lanka
C. yangbayensisTri & Onn, 2010[115]Yangbay bent-toed geckoVietnam
C. yathepyanensisGrismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[36]Yathe Pyan Cave bent-toed geckoMyanmar
C. yoshiiHikida, 1990Yoshi's bow-fingered geckoSabah, Borneo
C. ywanganensisGrismer, Wood, Thura, Quah, Murdoch, M. Grismer, Herr, Espinoza, & Lin, 2018Myanmar (Shan)
C. zebraicus(Taylor, 1962)Thailand
C. zhaoermiiShi & Zhao 2010 [116]Tibet, China
C. ziegleriNazarov, Orlov, Nguyen, & Ho, 2008 [117]Vietnam
C. zugiOliver, Tjaturadi, Mumpuni, Krey & Richards, 2008[118]Melanesia

Phylogeny

The following phylogeny is from Pyron, et al. (2013),[119] and includes 22 Cyrtodactylus species. Cyrtodactylus is a sister group of Hemidactylus.[119]

Cyrtodactylus 
 
 

Cyrtodactylus oldhami

 
 
 

Cyrtodactylus ayeyarwadyensis

 
 
 
 

Cyrtodactylus angularis

 
 

Cyrtodactylus jarujini

 
 
 
 
 

Cyrtodactylus triedrus

 
 
 
 

Cyrtodactylus marmoratus

 
 
 
 

Cyrtodactylus irregularis

 
 

Cyrtodactylus consobrinus

 
 
 
 

Cyrtodactylus annulatus

 
 
 

Cyrtodactylus philippinicus

 
 

Cyrtodactylus agusanensis

 
 
 
 
 
 
 
 
 

Cyrtodactylus intermedius

 
 
 

Cyrtodactylus pulchellus

 
 
 
 
 

Cyrtodactylus sermowaiensis

 
 

Cyrtodactylus loriae

 
 
 
 

Cyrtodactylus novaeguineae

 
 

Cyrtodactylus tuberculatus

 
 
 
 
 
 

Cyrtodactylus klugei

 
 

Cyrtodactylus robustus

 
 
 
 

Cyrtodactylus tripartitus

 
 
 

Cyrtodactylus epiroticus

 
 

Cyrtodactylus louisiadensis

 
 
 
 
 
 
 
 
 
 
 
 

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