Ruyangosaurus

Ruyangosaurus
Temporal range: Early Cretaceous, Aptian–Albian
Skeletal and life restorations
Scientific classification
Kingdom:Animalia
Phylum:Chordata
Clade:Dinosauria
Order:Saurischia
Suborder:Sauropodomorpha
Clade:Sauropoda
Clade:Titanosauriformes
Genus:Ruyangosaurus
Lu et al., 2009
Species

Ruyangosaurus (Ruyang County lizard) is a genus of titanosauriform sauropod dinosaur recovered from the Early Cretaceous Haoling Formation of China. The type species is R. giganteus, described in 2009 by a group of scientists from the Chinese Academy of Sciences and the Henan Geological Museum.[1] Along with Huanghetitan and Daxiatitan, Ruyangosaurus is among the largest dinosaurs discovered in Cretaceous Asia.

Classification

The describers of Ruyangosaurus assigned it to Andesauridae.[1] However, Andesauridae is not monophyletic and, as such, is no longer used.[2]

The most comprehensive phylogenetic dataset to include Ruyangosaurus finds it to be in a polytomy with Andesaurus and other basal titanosaurs and near-titanosaur somphospondyls, supporting the original description's assertion of phylogenetic proximity of it and Andesaurus, though not the precise taxonomic assignment.[3][4]

Macronaria

Tehuelchesaurus

Janenschia

Haestasaurus

Camarasaurus

Aragosaurus

Galveosaurus

Titanosauriformes

Brachiosauridae

Europasaurus

Vouivria

Brachiosaurus

Abydosaurus

Cedarosaurus

Giraffatitan

Lusotitan

Sonorasaurus

Venenosaurus

Somphospondyli

Euhelopodidae

Euhelopus

Erketu

Gobititan

Qiaowanlong

Phuwiangosaurus

Tangvayosaurus

Paluxysaurus

Sauroposeidon

Cloverly titanosauriform

Padillasaurus

Dongbeititan

Ligabuesaurus

Tastavinsaurus

Angolatitan

Chubutisaurus

Andesaurus

Huanghetitan liujiaxianensis

Huanghetitan ruyangensis

Ruyangosaurus

Wintonotitan

Baotianmansaurus

Dongyangosaurus

Savannasaurus

Diamantinasaurus

Daxiatitan

Xianshanosaurus

Lithostrotia

However, not all phylogenetic analyses have supported its position as a somphospondyl. A smaller phylogenetic dataset found Ruyangosaurus to be a non-titanosauriform macronarian based on additional material from the type locality.[5]

Macronaria

Camarasaurus

Europasaurus

Yunmenglong

Ruyangosaurus

Titanosauriformes

Brachiosauridae

Brachiosaurus

Giraffatitan

Abydosaurus

Paluxysaurus

Cedarosaurus

Somphospondyli

Euhelopus

Qiaowanlong

Erketu

Titanosauria

Andesaurus

Sonidosaurus

Bor Guvé titanosaur

Lithostrotia

Habitat

Ruyangosaurus shared its habitat with Xianshanosaurus, "Huanghetitan" ruyangensis, Yunmenglong, Luoyanggia, and Zhongyuansaurus. The type horizon of Ruyangosaurus was originally described as being of "early Late Cretaceous" age,[1] but recent work has assigned it an Aptian-Albian Age based on fieldwork and analysis of invertebrate and microfossil assemblages.[6]

See also

References

  1. 1 2 3 Lü J, Xu L, Jia S, Zhang X, Zhang J, Yang L, You H, Ji Q. (2009). "A new gigantic sauropod dinosaur from the Cretaceous of Ruyang, Henan, China". Geological Bulletin of China 28(1), 1-10.
  2. Mannion, Philip D.; Calvo, Jorge O. (2011). "Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albian-early Cenomanian) Río Limay Formation, Neuquén Province, Argentina: implications for titanosaur systematics". Zoological Journal of the Linnean Society. 163: 155–181. doi:10.1111/j.1096-3642.2011.00699.x.
  3. Mannion, Philip D.; Allain, Ronan; Moine, Olivier (2017). "The earliest known titanosauriform sauropod dinosaur and the evolution of Brachiosauridae". PeerJ. 5: e3217. doi:10.7717/peerj.3217.
  4. Royo-Torres, Rafael; Upchurch, Paul; Kirkland, James I.; DeBlieux, Donald D.; Foster, John R.; Cobos, Alberto; Alcalá, Luis (2017). "Descendants of the Jurassic turiasaurs from Iberia found refuge in the Early Cretaceous of western USA". Scientific Reports. 7: 14311. doi:10.1038/s41598-017-14677-2.
  5. Lü, Junchang; Pu, Hanyong; Xu, Li; Jia, Songhai; Zhang, Jiming; Shen, Caizhi (2014). Osteology of the giant sauropod dinosaur Ruyangosaurus giganteus Lü et al., 2009. Beijing: Geological Publishing House. ISBN 978-7-116-09074-3.
  6. Xu, L., Pan, Z.C., Wang, Z.H., Zhang, X.L., Jia, S.H., Lü, J.C., Jiang, B.L., 2012. Discovery and significance of the Cretaceous system in Ruyang Basin, Henan Province. Geological Review 58, 601-613.
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