Equisetopsida ''sensu lato''

Equisetopsida sensu lato; Temporal range: Late Silurian–Recent PreЄ Є O S D C P T J K Pg N
	Land plants
Scientific classification
Kingdom:	Plantae
(unranked):	Streptophyta
Class:	Equisetopsida; C. Agardh 1825
Subclasses
	About 13 subclasses. See text.
Synonyms
	Embryophyta

Equisetopsida is the name of a class of plants that traditionally contains the single genus Equisetum (horsestails). However, in 2009, in an article titled "A phylogenetic classification of the land plants to accompany APG III," Mark W. Chase and James L. Reveal proposed a much broader sense for the Equisetopsida class name. In their system, Equisetopsida includes all land plants (embryophytes). Their rationale was that "If the major clades of green algae are recognized as classes, then all land plants, the embryophytes, should be included in a single class..."^[1]

Subclasses

The class Equisetopsida sensu Chase & Reveal, 2009, contains the following subclasses. Traditional divisions and classes, along with common names are shown in parentheses.^[1]

bryophytes (non-vascular plants)
- Subclass Marchantiidae Engl. 1893 (div. Marchantiophyta, 3 classes - liverworts)
- Subclass Bryidae Engl. 1892 (div. Bryophyta, class Bryopsida - moss)
- Subclass Anthocerotidae Engl. 1893 (div. Anthocerotophyta, 2 classes - hornwort)
lycopodiophytes
- Subclass Lycopodiidae Bek. 1863 (div. Lycopodiophyta, 2 classes - clubmoss, spikemoss, quillworts)
monilophytes (ferns)^[2]
- Subclass Equisetidae Warm. 1883 (class Equisetopsida, s.s. - horsetails)
- Subclass Ophioglossidae Klinge 1882 (class Psilotopsida - whisk ferns, grape-ferns, moonworts)
- Subclass Marattiidae Klinge 1882 (class Marattiopsida - marattioid ferns)
- Subclass Polypodiidae Cronquist, Takht. & W. Zimm. 1966 (class Polypodiopsida - polypod ferns)
- Subclass Psilotidae Reveal 1996 [merged into Ophioglossidae by Christenhusz, 2011]
gymnosperms^[3]
- Subclass Cycadidae Pax 1894 (div. Cycadophyta, class Cycadopsida - cycads)
- Subclass Ginkgoidae Engl. 1897 (div. Ginkgophyta, class Ginkgoopsida, one extant genus: Ginko)
- Subclass Gnetidae Pax 1894 (div. Gnetophyta, class Gnetopsida - gnetophytes)
- Subclass Pinidae Cronquist, Takht. & W. Zimm. 1966 (div. Pinophyta, class Pinopsida - conifers)
angiosperms^[4]
- Subclass Magnoliidae Novák ex Takht. 1967 (Angiosperms - flowering plants)

Phylogeny

The following diagram shows a likely phylogenic relationship between the Equisetopsida subclasses.

Equisetopsida

	Marchantiidae

	Bryidae

	Anthocerotidae

	Lycopodiidae

	Equisetidae

	Ophioglossidae

	Marattiidae

	Polypodiidae

	Cycadidae

	Ginkgoidae

	Gnetidae

	Pinidae

	Magnoliidae

References

1 2 Mark W. Chase; James L. Reveal (2009). "A phylogenetic classification of the land plants to accompany APG III". Botanical Journal of the Linnean Society. 161: 122–127. doi:10.1111/j.1095-8339.2009.01002.x.
↑ Maarten J. M. Christenhusz; Xian-Chun Zhang; Harald Schneider (2011). "A linear sequence of extant families and genera of lycophytes and ferns" (PDF). Phytotaxa. 19: 7–54.
↑ Maarten J. M. Christenhusz; James L. Reveal; Aljos Farjon; Martin F. Gardner; Robert R. Mill; Mark W. Chase (2011). "A new classification and linear sequence of extant gymnosperms" (PDF). Phytotaxa. 19: 55–70.
↑ James L. Reveal; Mark W. Chase (2011). "APG III: Bibliographical Information and Synonymy of Magnoliidae" (PDF). Phytotaxa. 19: 71–134.

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[Chase-2009-1] 1 2 Mark W. Chase; James L. Reveal (2009). "A phylogenetic classification of the land plants to accompany APG III". Botanical Journal of the Linnean Society. 161: 122–127. doi:10.1111/j.1095-8339.2009.01002.x.

[Christenhusz-2011f-2] Maarten J. M. Christenhusz; Xian-Chun Zhang; Harald Schneider (2011). "A linear sequence of extant families and genera of lycophytes and ferns" (PDF). Phytotaxa. 19: 7–54.

[Christenhusz-2011g-3] Maarten J. M. Christenhusz; James L. Reveal; Aljos Farjon; Martin F. Gardner; Robert R. Mill; Mark W. Chase (2011). "A new classification and linear sequence of extant gymnosperms" (PDF). Phytotaxa. 19: 55–70.

[Reveal-2011-4] James L. Reveal; Mark W. Chase (2011). "APG III: Bibliographical Information and Synonymy of Magnoliidae" (PDF). Phytotaxa. 19: 71–134.