Phengodidae

Phengodidae
	Phengodes orbignyi
	Taxonomische indeling
Rijk:	Animalia (Dieren)
Stam:	Arthropoda (Geleedpotigen)
Klasse:	Insecta (Insecten)
Orde:	Coleoptera (Kevers)
Onderorde:	Polyphaga
Infraorde:	Elateriformia
Superfamilie:	Elateroidea
	Familie
	Phengodidae; LeConte, 1861
	Afbeeldingen op Wikimedia Commons
	Phengodidae op Wikispecies
Portaal	Biologie ; Insecten

Phengodidae zijn een familie van kevers. De familie is voor het eerst wetenschappelijk beschreven in 1861 door LeConte. Het is een van de families van glimwormen.

Taxonomie

De volgende taxa zijn bij de familie ingedeeld:[1]

Onderfamilie Phengodinae LeConte, 1861
- = Pseudophengodidae Pic, 1930
- Geslacht Microphengodes Wittmer, 1976
  - Microphengodes howdeni Wittmer, 1976
  - Microphengodes longicornis Wittmer, 1976
- Geslacht Phengodes Illiger, 1807
  - = Astraptor Murray, 1868
  - Ondergeslacht Phengodella Wittmer
    - Phengodes atezcanus Zaragoza, 1981
    - Phengodes bella Barber, 1913
    - Phengodes bimaculata Gorham, 1881
      - = Phengodes nevermani Pic, 1937
      - = Phengodes notaticollis Pic, 1937:
    - Phengodes bipennifera Gorham, 1881
      - = Phengodes brasiliensis Pic, 1925:
    - Phengodes brailovskyi Zaragoza & Wittmer, 1986
    - Phengodes championi Pic, 1927
    - Phengodes ecuadoriana Witmer, 1988
    - Phengodes fenestrata Wittmer, 1975
    - Phengodes frontalis LeConte, 1881
    - Phengodes insignis Bourgeois, 1888
    - Phengodes insulcata Pic, 1925
      - = Phengodes atricolor Pic, 1937
    - Phengodes leonilae Zaragoza & Wittmer, 1986
    - Phengodes minor Gorham, 1881
    - Phengodes nigricornis Gorham, 1881
      - = Phengodes bipennifera Gorham, 1881
      - = Phengodes gorhami Pic, 1927
    - Phengodes tuxtlaensis Zaragoza, 1989
    - Phengodes varicolor Zaragoza & Wittmer, 1986
    - Phengodes variicornis Zaragoza & Wittmer, 1986
    - Phengodes venezolana Wittmer, 1988
  - Ondergeslacht Phengodes Illiger, 1807
    - Phengodes arizonensis Wittmer, 1975
    - Phengodes bolivari Zarazoga, 1980
    - Phengodes fusciceps LeConte, 1861
    - Phengodes inflata Wittmer, 1975
    - Phengodes laticollis LeConte, 1881
    - Phengodes mexicana Wittmer, 1975
    - Phengodes nigromaculata Wittmer, 1975
    - Phengodes plumosa (Oliver, 1790)
      - = Lampyris plumosa Oliver, 1790
    - Phengodes vazquezae Zarazoga, 1978
- Geslacht Pseudophengodes Pic, 1930
  - Pseudophengodes bordoni Wittmer, 1988
  - Pseudophengodes brasiliensis Wittmer, 1976
  - Pseudophengodes cincinnata (Erichson, 1847)
    - = Phengodes cincinnata Erichson, 1847
  - Pseudophengodes equatoriana Wittmer, 1976
  - seudophengodes fernandezi Wittmer, 1976
  - Pseudophengodes flavicollis (Leach, 1824)
    - = Phengodes flavicollis Leach, 1824
  - Pseudophengodes floccosa (Erichson, 1847)
    - = Phengodes floccosa Erichson, 1847
  - Pseudophengodes foveiocollis Wittmer, 1976
  - Pseudophengodes fusca (Gorham, 1881)
    - = Phengodes fusca Gorham, 1881
  - Pseudophengodes gracilicornis Wittmer, 1976
  - Pseudophengodes marginata (Pic, 1925)
    - = Phengodes marginata Pic, 1925
  - Pseudophengodes marmoratus Wittmer, 1996
  - Pseudophengodes meridiana Wittmer, 1976
  - Pseudophengodes notaticollis (Pic, 1925)
    - = Phengodes notaticollis Pic, 1925
  - Pseudophengodes onorei Wittmer, 1996
  - Pseudophengodes orbignyi (Blanchard, 1837)
    - = Phengodes orbignyi Blanchard, 1837
  - Pseudophengodes penai Wittmer, 1976
  - Pseudophengodes polita Wittmer, 1976
  - Pseudophengodes pulchella (Guerin, 1843)
    - = Phengodes pulchella Guerin, 1843
  - Pseudophengodes puncticollis Wittmer, 1976
  - Pseudophengodes roulini (Guerin, 1843)
    - = Phengodes roulini Guerin, 1843
  - Pseudophengodes ruficollis (Motschulsky, 1854)
    - = Phengodes ruficollis Motschulsky, 1854
- Geslacht Zarhipis J.LeConte, 1880
  - = Zarhippus Woodworth, 1913
  - = Zarrhipis Fall, 1923
  - Zarhipis integripennis (LeConte, 1874)
    - = Phengodes integripennis LeConte, 1874
    - = Zarrhipis alamedae Fall, 1923
    - = Zarrhipis amictus Fall, 1923
    - = Zarrhipis brevicollis Fall, 1923
    - = Zarhipis piciventris LeConte, 1881
    - = Zarhipis riversi Horn, 1885
    - = Zarhipis ruficollis LeConte, 1881
  - Zarhipis tiemanni Linsdale, 1964
  - Zarhipis truncaticeps (Fall, 1923
    - = Zarrhipis truncaticeps Fall, 1923
Onderfamilie Mastinocerinae LeConte, 1881
- Geslacht Akamboja Roza, Quintino, Mermudes & Silveira, 2017[2]
  - Akamboja caparaoensis Roza, Quintino, Mermudes & Silveira, 2017[3]
  - Akamboja cleidae Roza, Quintino, Mermudes & Silveira, 2017[4]
  - Akamboja minimum Roza, Quintino, Mermudes & Silveira, 2017[5]
  - Akamboja monteirorum Roza, Quintino, Mermudes & Silveira, 2017[6]
  - Akamboja tenebrae Roza, Quintino, Mermudes & Silveira, 2017[7]
- Geslacht Baeoscelis Spinola, 1854
  - Baeoscelis osculata Spinola, 1854
- Geslacht Brasilocerus Wittmer, 1963
  - = Brasibcerus
  - Brasilocerus belemensis Wittmer, 1988
  - Brasilocerus dietrichi (Wittmer, 1963)
    - = Mastinocerus dietrichi Wittmer, 1963
  - Brasilocerus espiritensis Wittmer, 1963
  - Brasilocerus minasensis Wittmer, 1976
  - Brasilocerus nigerrimus Wittmer, 1996
  - Brasilocerus nigrofasciatus (Pic, 1915)
    - = Phrixothrix nigrofasciatus Pic, 1915
  - Brasilocerus oberthuri (Pic, 1955)
    - = Phrixothrix oberthuri Pic, 1955
    - = Brasilocerus impressicollis Wittmer, 1970
  - Brasilocerus opacus (Pic, 1937)
    - = Phrixothrix opacus Pic, 1937
    - = Mastinocerus callanganus Wittmer, 1963
    - = Phrixothrix subopacus Pic, 1954
  - Brasilocerus wygodzinsyi Wittmer, 1976
- Geslacht Cenophengus J.LeConte, 1881
  - Cenophengus breviplumatus Wittmer, 1976
  - Cenophengus brunneus Wittmer, 1976
  - Cenophengus ciceroi Wittmer, 1981
  - Cenophengus debilis .LeConte, 1881
  - Cenophengus gorhami Zarazoga, 1986
  - Cenophengus guerrerensis Zaragoza, 1991
  - Cenophengus howdeni Zarazoga, 1986
  - Cenophengus longicollis Wittmer, 1976
  - Cenophengus magnus Zaragoza, 1988
  - Cenophengus major Wittmer, 1976
  - Cenophengus marmoratus
  - Cenophengus pallidus Schaeffer, 1904
  - Cenophengus pedregalensis Zaragoza, 1975
  - Cenophengus punctatissimus Wittmer, 1976
  - Cenophengus villae Zaragoza-Caballero, 1984
  - Cenophengus wittmeri Zaragoza-Caballero, 1984
- Geslacht Cephalophrixothrix Wittmer, 1976
  - Cephalophrixothrix clypeatus Wittmer, 1976
  - Cephalophrixothrix columbianus Wittmer, 1976
  - Cephalophrixothrix nigerrimus Wittmer, 1976
- Geslacht Decamastinocerus Wittmer, 1988
  - Decamastinocerus parvimandibularis Wittmer, 1988
- Geslacht Distremocephalus Wittmer, 1976
  - Distremocephalus barrerai Zarazoga, 1986
  - Distremocephalus beutelspacheri Zarazoga, 1986
  - Distremocephalus buenoi Zarazoga, 1986
  - Distremocephalus californicus (Van Dyke, 1918)
    - = Mastinocerus californicus Van Dyke, 1918
  - Distremocephalus chiapensis Zarazoga, 1986
  - Distremocephalus leonilae Zarazoga, 1986
  - Distremocephalus mexicanus (Wittmer, 1963)
    - = Mastinocerus mexicanus Wittmer, 1963
  - Distremocephalus opaculus (Horn, 1895)
    - = Mastinocerus opaculus Horn, 1895
  - Distremocephalus rufocaudatus Zarazoga, 1986
  - Distremocephalus texanus (LeConte, 1874)
    - = Mastinocerus texanus LeConte, 1874
  - Distremocephalus wittmeri Zarazoga, 1986
- Geslacht Eurymastinocerus Wittmer, 1976
  - Eurymastinocerus arawakensis Wittmer, 1976
  - Eurymastinocerus columbiana (Wittmer, 1963)
    - = Euryopa columbiana Wittmer, 1963
  - Eurymastinocerus niger Gorham, 1881
    - = Euryopa nigra Gorham, 1881
  - Eurymastinocerus politicollis Wittmer, 1988
  - Eurymastinocerus reductipennis (Wittmer, 1970
    - = Mastinocerus reductipennis Wittmer, 1970
- Geslacht Euryognathus Wittmer, 1976
  - Euryognathus bicoloratus Wittmer, 1976
  - Euryognathus venezolanus (Wittmer, 1956)
    - = Mastinocerus venezolanus Wittmer, 1956
- Geslacht Euryopa Gorham, 1881
  - = Bruchodrilus Pic, 1926
  - Euryopa brasiliensis Wittmer, 1976
  - Euryopa caucaensis Wittmer, 1976
  - Euryopa clarindae Wittmer, 1996
  - Euryopa columbiana Wittmer, 1963
  - Euryopa laurae Wittmer, 1996
  - Euryopa nigra Gorham, 1881
  - Euryopa opacipennis (Pic, 1926)
    - = Bruchodrilus opacipennis Pic, 1926
    - = Euryopa robustior Pic, 1929
  - Euryopa peckorum Wittmer, 1996
  - Euryopa ruficeps Wittmer, 1976
  - Euryopa singularis Gorham, 1881
- Geslacht Howdenia Wittmer, 1976
  - Howdenia coroicoensis Wittmer, 1988
  - Howdenia fischeri (Pic, 1937)
    - = Phrixothrix fischeri Pic, 1937
  - Howdenia frontalis (Wittmer, 1963)
    - = Taximastinocerus frontalis Wittmer, 1963
  - Howdenia golbachi Wittmer, 1988
  - Howdenia minutissima Wittmer, 1988
  - Howdenia neblinensis Wittmer, 1988
  - Howdenia nigerrima Wittmer, 1976
  - Howdenia punctata Wittmer, 1976
  - Howdenia robusta Wittmer, 1988
- Geslacht Mastinocerus Solier in Gay, 1849
  - Ondergeslacht Mastinocerus Solier in Gay, 1849
    - = Mastigocerus Gemminger & Harold, 1869
    - Mastinocerus araucanus Wittmer, 1963
    - Mastinocerus atriceps (Pic, 1915)
      - = Phrixothrix atriceps Pic, 1915
      - = Phrixothrix argentinus Pic, 1915
    - Mastinocerus brevipennis Solier in Gay, 1849
    - Mastinocerus chilensis Wittmer, 1976
    - Mastinocerus germaini Pic, 1930
    - Mastinocerus kuscheli Wittmer, 1976
    - Mastinocerus nigeropacus Wittmer, 1970
    - Mastinocerus nigriceps Wittmer, 1963
    - Mastinocerus nigricollis (Pic, 1915)
      - = Phrixothrix nigricollis Pic, 1915
    - Mastinocerus nigroapicalis Pic, 1915
      - = Phrixothrix nigroapicalis
    - Mastinocerus obscurus Wittmer, 1976
    - Mastinocerus patruelis (Pic, 1915)
      - = Phrixothrix patruelis Pic, 1915
    - Mastinocerus punctatus Wittmer, 1963
    - Mastinocerus ramosus (Gorham, 1881)
      - = Ptorthodius ramosus Gorham, 1881
    - Mastinocerus ruficollis Wittmer, 1976
    - Mastinocerus travassosi Wittmer, 1963
    - Mastinocerus uruguayensis (Berg, 1886
      - = Phengodes uruguayensis Berg, 1886
  - Ondergeslacht Mastinocerus incertae
    - Mastinocerus callanganus Wittmer, 1963
  - Ondergeslacht Paramastinocerus Wittmer, 1976
    - Mastinocerus janeirensis Wittmer, 1976
    - Mastinocerus ondulatus Wittmer, 1963
    - Mastinocerus pauloensis Wittmer, 1963
    - Mastinocerus peruanus (Wittmer, 1976)
      - = Matinocerus peruanus Wittmer, 1976
- Geslacht Mastinomorphus Wittmer, 1976
  - Mastinomorphus atacamensis (Wittmer, 1963)
    - = Mastinocerus atacamensis Wittmer, 1963
  - Mastinomorphus impressiceps Wittmer, 1997
  - Mastinomorphus martinezi (Wittmer, 1963)
    - = Mastinocerus martinezi Wittmer, 1963
  - Mastinomorphus minutus Wittmer, 1976
  - Mastinomorphus misionensis (Wittmer, 1950)
    - = Phrixothrix misionensis Wittmer, 1950
  - Mastinomorphus obscuripennis Wittmer, 1976
  - Mastinomorphus pampaensis (Wittmer, 1950)
    - = Phrixothrix pampaensis Wittmer, 1950
  - Mastinomorphus piceipennis (Pic, 1929)
    - = Phrixothrix piceipennis Pic, 1929
  - Mastinomorphus rufescens (Pic, 1938)
    - = Phrixothrix rufescens Pic, 1938
  - Mastinomorphus ruficeps (Pic, 1926)
    - = Phrixothrix ruficeps Pic, 1926
  - Mastinomorphus vicunaensis Wittmer, 1976
  - Mastinomorphus weiseri (Pic, 1926)
    - = Phrixothrix weiseri Pic, 1926
- Geslacht Mastinowittmerus Zaragoza, 1984
  - Mastinowittmerus mexicanus Zaragoza, 1984
  - Mastinowittmerus vazquezae Zarazoga, 1986
- Geslacht Neophengus Wittmer, 1976
  - Neophengus chilensis Wittmer, 1976
  - Neophengus huantaensis Wittmer, 1976
  - Neophengus nanus (Wittmer, 1948)
    - = Phrixothrix nanus Wittmer, 1948
  - Neophengus penai (Wittmer, 1963)
    - = Cenophengus penai Wittmer, 1963
- Geslacht Nephromma Wittmer, 1976
  - Nephromma alvarengai Wittmer, 1996
  - Nephromma barberi (Wittmer, 1976)
    - = Nephroma barberi Wittmer, 1976
- Geslacht Oxymastinocerus Wittmer, 1963
  - Oxymastinocerus bridarolli Wittmer, 1963
  - Oxymastinocerus fulvus (Philippi, 1864)
    - = Mastinocerus fulvus Philippi, 1864
  - Oxymastinocerus nigripennis Wittmer, 1988
  - Oxymastinocerus pecki Wittmer, 1976
  - Oxymastinocerus peruanus (Wittmer, 1988)
    - = Mastinocerus peruanus Wittmer, 1956
  - Oxymastinocerus rufotestaceus Wittmer, 1976
  - Oxymastinocerus unicolor (Pic, 1926)
    - = Phrixothrix unicolor Pic, 1926
  - Oxymastinocerus venezolanus Wittmer, 1976
- Geslacht Paraphrixothrix Zaragoza-Caballero, 2010[8]
  - Paraphrixothrix ecuadoranus Zaragoza-Caballero, 2010
- Geslacht Paraptorthodius Schaeffer, 1904
  - Paraptorthodius mirabilis Schaeffer, 1904
  - Paraptorthodius schaefferi (Zaragoza, 1989:88)
    - = Paratorthodius schaefferi Zaragoza, 1989
- Geslacht Phrixothrix Olivier, 1909
  - Phrixothrix acuminatus Pic, 1929
  - Phrixothrix alboterminatus Wittmer, 1963
  - Phrixothrix belemensis Wittmer, 1976
  - Phrixothrix clypeatus Wittmer, 1992
  - Phrixothrix gibbosus Wittmer, 1976
  - Phrixothrix heydeni Olivier, 1910
  - Phrixothrix hieronymi (Haase, 1886)
    - = Phengodes hieronymi Haase, 1886
    - = Phrixothrix obscuripes Pic, 1915
  - Phrixothrix hirtus Olivier, 1909
  - Phrixothrix microphthalmus Wittmer, 1976
  - Phrixothrix obscurus Pic, 1915
  - Phrixothrix peruanus Wittmer, 1963
  - Phrixothrix pickeli Pic, 1933
  - Phrixothrix reducticornis Wittmer, 1963
  - Phrixothrix staphylinoides Wittmer, 1963
  - Phrixothrix tiemanni Wittmer, 1970
  - Phrixothrix vianai Wittmer, 1988
  - Phrixothrix vivianii Wittmer, 1992
- Geslacht Pseudomastinocerus Wittmer, 1963
  - Pseudomastinocerus chiriquiensis Wittmer, 1963
  - Pseudomastinocerus costaricanus Wittmer, 1988
  - Pseudomastinocerus freyi Wittmer, 1963
  - Pseudomastinocerus laticeps (Pic, 1937)
    - = Phrixothrix laticeps Pic, 1937
  - Pseudomastinocerus panamensis Wittmer, 1976
  - Pseudomastinocerus ruficeps (Pic, 1925)
    - = Phengodes ruficeps Pic, 1925
  - Pseudomastinocerus tinalandicus Wittmer, 1976
- Geslacht Ptorthodiellus Wittmer, 1976
  - Ptorthodiellus araguaicus Wittmer, 1976
  - Ptorthodiellus trinidadicus Wittmer, 1976
- Geslacht Ptorthodius Gorham, 1881
  - - = Cephalodrilus Pic
  - Ptorthodius atricornis (Pic, 1929)
    - = Phrixothrix atricornis Pic, 1929
  - Ptorthodius lanei Wittmer, 1963[9]
  - Ptorthodius mandibularis Gorham, 1881
    - = Cephalodrilus atrosignatus Pic
  - Ptorthodius quiguasensis Wittmer, 1996
- Geslacht Spangleriella Wittmer, 1988
  - Spangleriella vittata Wittmer, 1988
- Geslacht Stenophrixothrix Wittmer, 1963
  - Stenophrixothrix bogotensis (Pic, 1925)
    - = Phengodes bogotensis Pic, 1925
  - Stenophrixothrix chiriquensis Wittmer, 1988
  - Stenophrixothrix curticollis (Pic, 1925)
    - = Phengodes curticollis Pic, 1925
  - Stenophrixothrix darlingtoni Wittmer, 1976
  - Stenophrixothrix ecuadoranus Wittmer, 1988
  - Stenophrixothrix elongatus Wittmer, 1988
  - Stenophrixothrix espiritensis Wittmer, 1996
  - Stenophrixothrix fuscus Gorham, 1881
    - = Euryopa fusca Gorham, 1881
  - Stenophrixothrix howdeni Wittmer, 1988
  - Stenophrixothrix longipes Wittmer, 1976
  - Stenophrixothrix mandibularis Wittmer, 1988
  - Stenophrixothrix martinezi Wittmer, 1988
  - Stenophrixothrix minor Wittmer, 1976
  - Stenophrixothrix montanus Wittmer, 1976
  - Stenophrixothrix nigripennis Wittmer, 1976
  - Stenophrixothrix oculatus Wittmer, 1988
  - Stenophrixothrix pallens (Berg, 1885:232)
    - = Phengodes pallens Berg, 1885
  - Stenophrixothrix panamensis Wittmer, 1988
  - Stenophrixothrix pseudomandibularis Wittmer, 1988
- Geslacht Taximastinocerus Wittmer, 1963
  - Taximastinocerus beniensis Wittmer, 1976
  - Taximastinocerus brasiliensis Wittmer, 1976
  - Taximastinocerus breviplumatus Wittmer, 1988
  - Taximastinocerus brunneus Gorham, 1881
    - = Euryopa brunea Gorham, 1881
  - Taximastinocerus cephalotes (Pic, 1938)
    - = Phrixothrix cephalotes Pic, 1938
  - Taximastinocerus frontalis Wittmer, 1963
  - Taximastinocerus hermanni Wittmer, 1997
  - Taximastinocerus hickeri (Pic, 1929)
    - = Phrixothrix hickeri Pic, 1929
  - Taximastinocerus kissingeri Wittmer, 1963
  - Taximastinocerus levefumidus Wittmer, 1988
  - Taximastinocerus neblinensis Wittmer, 1988
  - Taximastinocerus nigricolor Wittmer, 1988
  - Taximastinocerus pallidus (Pic, 1938)
    - = Phrixothrix pallidus Pic, 1938
  - Taximastinocerus paralleleus Wittmer, 1976
  - Taximastinocerus plaumanni Wittmer, 1963
  - Taximastinocerus pseudobrunneus Wittmer, 1988
  - Taximastinocerus puncticollis Wittmer, 1976
Onderfamilie Penicillophorinae Paulus, 1975
- Geslacht Acladocera Wittmer, 1981
  - Acladocera hispaniolae Wittmer, 1981
- Geslacht Adendrocera Wittmer, 1976
  - Adendrocera flavulum (Wittmer, 1976)
    - = Anendocera flavulum Wittmer, 1976
- Geslacht Penicillophorus Paulus, 1974
  - Penicillophorus ctenotarsus Paulus, 1974
- Geslacht Tarsakanthos Zaragoza-Caballero, 2008[10]
  - Tarsakanthos minuta Zaragoza-Caballero, 2008[10]
- Geslacht Walterius Zaragoza-Caballero, 2008[10]
  - Walterius caballeroae Zaragoza-Caballero, 2008[10]
  - Walterius emilioi Zaragoza-Caballero, 2008[10]

Bronnen, noten en/of referenties

V.w.b. onderfamilies Bouchard, P. et al (2011) Family-group names in Coleoptera (Insecta). ZooKeys, 88: 1-972. DOI:10.3897/zookeys.88.807 ISBN 978-954-642-583-6 (hardback) ISBN 978-954-642-584-3 (paperback) ZooBank
Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 503-504
Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 516-518
Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523
Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 505-515
Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 515
Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 516
Zaragoza-Caballero, S. (2010) Paraphrixothrix ecuadoranus, a new genus and species of Phengodidae (Coleoptera: Mastinocerinae) from Ecuador. Pan-Pacific entomologist, 86(4): 105–110. DOI:10.3956/2007-40.1
Wittmer, W. (1963) Zur Kenntnis der Phengodidae (Coleoptera) (25. Beitrag zuer Kenntnis der neotropischen Malacodermata). Mitteilungen der Schweizerischen entomologischen Gesellschaft, 36, 73–99.
Zaragoza-Caballero, S. (2008) Two new genera and three new species of Penicillophorinae (Coleoptera: Phengodidae) of Mexico. Revista mexicana de biodiversidad, 79(2): 363–368. PDF

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[1] V.w.b. onderfamilies Bouchard, P. et al (2011) Family-group names in Coleoptera (Insecta). ZooKeys, 88: 1-972. DOI:10.3897/zookeys.88.807 ISBN 978-954-642-583-6 (hardback) ISBN 978-954-642-584-3 (paperback) ZooBank

[2] Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 503-504

[3] Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 516-518

[4] Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523

[5] Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 505-515

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[Zaragoza2008-10] Zaragoza-Caballero, S. (2008) Two new genera and three new species of Penicillophorinae (Coleoptera: Phengodidae) of Mexico. Revista mexicana de biodiversidad, 79(2): 363–368. PDF