Kimmeridge Clay

Kimmeridge Clay
Stratigraphic range: Kimmeridgian-Tithonian
Kimmeridge Clay; Stratigraphic range: Kimmeridgian-Tithonian
	Grey cliffs of Upper Kimmeridge Clay above the beach at Egmont Bight
Type	Geological formation
Unit of	Ancholme Group
Sub-units	Birch Sandstone Member, Burns Sandstone Member, Claymore Sandstone Member, Dirk Sandstone Member, Magnus Sandstone Member, Ptarmigan Sandstone Member, Ribble Sandstone Member
Underlies	Portland Sandstone Formation
Overlies	Ampthill Clay, Corallian Group
Lithology
Primary	Mudstone
Other	Siltstone, Sandstone, Conglomerate
Location
Region	England (surface); North Sea (subsurface)
Country	England
Type section
Named for	Kimmeridge Bay

The Kimmeridge Clay is a sedimentary deposit of fossiliferous marine clay which is of Jurassic age and occurs in Europe. This rock formation is the major source rock for North Sea oil. The fossil fauna of the Kimmeridge Clay includes turtles, crocodiles, sauropods, plesiosaurs, pliosaurs and ichthyosaurs, as well as a number of invertebrate species.

Description

Kimmeridge Clay is named after the village of Kimmeridge on the Dorset coast of England, where it is well exposed and forms part of the Jurassic Coast World Heritage Site. It exists across England, in a band stretching from Dorset in the south-west, north-east to East Anglia.

The Humber Bridge's foundations are in the Kimmeridge Clay deposits under the Humber estuary.

Economic importance

Kimmeridge Clay is arguably the most economically important unit of rocks in the whole of Europe, being the major source rock for oil fields in the North Sea hydrocarbon province. It has distinctive physical properties, log responses, and palynological signature.

A Kimmeridge Oil Shale Project (KOSP) has been pursued by the UK based oil and shale gas exploration company Cuadrilla Resources.^[1]^:6

Vertebrate fauna

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

Fauna uncovered from the Kimmeridge Clay include:^[2]

Ray-finned fish

Ray-finned fishes of the Kimmeridge clay Formation
Genus	Species	Location	Stratigraphic Position	Abundance	Notes	Images
Thrissops^[3]^[4]	Indeterminate	Dorset		Most common Kimmeridge clay fish, known from several complete specimens	Housed at the etches collection, discovered by Steve Etches	Complete specimen
Pachythrissops	Indeterminate	Dorset			Housed at the etches collection, discovered by Steve Etches
Allothrissops	Indeterminate	Dorset			Housed at the etches collection, discovered by Steve Etches
Chondrostei	Indeterminate	Dorset		One specimen, a fin	Housed at the etches collection, discovered by Steve Etches
Lepidotes	Indeterminate	Dorset			Housed at the etches collection, discovered by Steve Etches
Gyrodus	Indeterminate	Dorset			Housed at the etches collection, discovered by Steve Etches
Caturus	Indeterminate	Dorset			Housed at the etches collection, discovered by Steve Etches
Aspidorhynchus	Indeterminate	Dorset			Housed at the etches collection, discovered by Steve Etches
Hypsocormus	H.tenuirostris	Dorset		Rare	Housed at the etches collection, discovered by Steve Etches
Pachycormus	Indeterminate	Dorset		Rare	Housed at the etches collection, discovered by Steve Etches
Eurycormus	Indeterminate	Dorset		Rare, one complete specimen	Housed at the etches collection, discovered by Steve Etches
Leptolepidae	Indeterminate	Dorset		Fairly common, multiple near complete specimens.	Housed at the etches collection, discovered by Steve Etches

Lobe-finned fish

Lobe-finned fishes of the Kimmeridge clay Formation
Genus	Species	Location	Stratigraphic Position	Abundance	Notes	Images
Holophagus	Indeterminate	Dorset		One specimen, cranial material	Housed at the etches collection, discovered by Steve Etches. More than 2 metres long
Coelocanthus	Indeterminate	Dorset		One specimen, a portion of the head	Housed at the etches collection, discovered by Steve Etches

Cartilaginous fish

Cartilaginous fishes of the Kimmeridge clay Formation
Genus	Species	Location	Stratigraphic Position	Abundance	Notes	Images
Asteracanthus	Indeterminate	Dorset		Known from many dorsal spines	Housed at the etches collection, discovered by Steve Etches
Chimaera	Indeterminate	Dorset		Known from many dorsal spines	Housed at the etches collection, discovered by Steve Etches
Hybodus	Indeterminate	Dorset		Known from many dorsal spines, perhaps a complete head	Housed at the etches collection, discovered by Steve Etches
Ischyodus	Indeterminate	Dorset		One specimen	Housed at the etches collection, discovered by Steve Etches
Rhinobatidae	Indeterminate	Dorset		Known from a complete specimen, and other isolated remains	Housed at the etches collection, discovered by Steve Etches

Archosaurs

Thalattosuchians

Genus	Species	Notes
Cricosaurus	C. gracilis	A metriorhynchine metriorhynchid
Dakosaurus	D. maximus	A geosaurine metriorhynchid
Metriorhynchus	M. geoffreyi	A metriorhynchine metriorhynchid
Plesiosuchus	P. manseli	A geosaurine metriorhynchid
Steneosaurus	S. megarhinus	A teleosaurid
Torvoneustes^[5]^[6]	T. carpenteri^[5]	A geosaurine metriorhynchid
Torvoneustes^[5]^[6]	T. coryphaeus^[7]	A geosaurine metriorhynchid

Ornithischians

Indeterminate nodosaurid remains have been found in Wiltshire, England.^[2] Indeterminate stegosaurid remains have been found in Dorset and Wiltshire, England.^[2]

Genus	Species	Location	Member	Abundance	Notes	Images
Cumnoria^[2]	C. prestwichii^[2]	Oxfordshire^[2]		"Fragmentary skull and skeleton."^[8]
Dacentrurus^[2]	D. armatus^[2]	Cambridgeshire^[2] Dorset^[2] Wiltshire^[2]			Wiltshire remains include specimens previously referred to Omosaurus armatus and O. hastiger.^[2]	Dacentrurus
Ornithopoda	Indeterminate^[2]	Dorset?^[2]			Kimmeridge clay remains considered to represent a possible close relative of Bugenasaura^[9] are now regarded as the remains of an indeterminate euornithopod.^[2](The specimen may have had its locality and horizon mislabelled.)
	Omosaurus^[2]	O.armatus^[2]	Wiltshire^[2]			Reclassified as Dacentrurus armatus because the generic name Omosaurus was preoccupied.^[2]
O. hastiger^[2]	Omosaurus^[2]	Wiltshire^[2]

Saurischians

Indeterminate ornithomimmid remains have been found in Dorset, England.^[2] An undescribed theropod genus was found in Dorset.^[2]

Genus	Species	Location	Material	Notes
Bothriospondylus^[2]	B. suffosus^[2]	Wiltshire^[2]	"[Seven] dorsal and sacral centra."^[10]
Cetiosaurus^[2]	C. humerocristatus^[2]	Dorset^[2]		Now Duriatitan.^[11]
Cetiosaurus^[2]	Indeterminate^[2]	Oxfordshire^[2]		Remains previously referred to an indeterminate species of Cetiosaurus are now regarded as indeterminate sauropod material.^[2]
Duriatitan	D. humerocristatus	Dorset^[2]	Humerus^[12]	A titanosauriform^[11]
Gigantosaurus^[2]	G. megalonyx^[2]	Cambridgeshire^[2]		Remains previously referred to Gigantosaurus megalonyx are now regarded as indeterminate sauropod material.^[2]
Ischyrosaurus^[2]	I. manseli^[2]	Oxfordshire^[2]	"Humerus."^[13]	Remains previously referred to Ischyrosaurus manseli are now regarded as indeterminate sauropod material.^[2]
Juratyrant^[14]^[15]	J. langhami	Dorset	Partial skeleton	A tyrannosaur
Theropoda^[2]	Indeterminate	Wiltshire^[2] Dorset^[2]		Remains previously referred to Megalosaurus are now regarded as indeterminate theropod material.^[2]
Sauropoda^[2]	Indeterminate^[2]	Oxfordshire^[2] Cambridgeshire^[2] Norfolk^[2]		Remains previously attributed to one or more indeterminate species of Ornithopsis (incl. O. leedsii) are now regarded as possible indeterminate sauropod material.^[2]

Pterosaurs

Genus	Species	location	Material	notes
Cuspicephalus^[16]	C. scarfi	Dorset	Partial Skull	Missing Crest, lower jaw and dentition
Rhamphorhynchus	R.etchesi	Dorset
Germanodactylus	indeterminate	Dorset

Plesiosaurs

Genus	Species	Abundance	Notes
Bathyspondylus	B. swindoniensis		Plesiosaur of unknown affinities
Colymbosaurus	C. megadeirus		A cryptoclidid
Colymbosaurus	C. trochantericus		Nomen dubium
Kimmerosaurus	K. langhami		A cryptoclidid
Plesiosaurus	"P." manseli		Distinct from Colymbosaurus.^[17]
Pliosaurus^[18]^[19]	P. brachydeirus		A thalassophonean pliosaurid
	P. brachyspondylus
	P. carpenteri		A thalassophonean pliosaurid
	P. kevani		A thalassophonean pliosaurid
	P. portentificus^[20]		A thalassophonean pliosaurid
	P. ?rossicus		A thalassophonean pliosaurid; taxonomic identification of specimens tentative^[18]
	P. westburyensis		A thalassophonean pliosaurid
	P. sp. 1	Partial skeleton, CAMSM J.35991	A thalassophonean pliosaurid; previously assigned to the nomen dubium P. brachyspondylus^[18]^[19]
	P. sp. 2	Mandible, NHMUK PV OR 39362	A thalassophonean pliosaurid; previously assigned to the nomen dubium P. macromerus^[18]^[19]
Spitrasaurus	Indeterminate

Ichthyosaurs

Ray-finned fishes of the Kimmeridge clay Formation
Genus	Species	Location	Stratigraphic Position	Abundance	Notes	Images
Brachypterygius	B.extremus	Dorset
Ichthyosauridae	Indeterminate	Dorset			Giant, near complete specimen. Proposed to have been 6 metres long when complete. Housed at the Etches collection in dorset.
Macropterygius	M.trigonus	Dorset			Nomen dubium - classified by a single vertebra
Nannopterygius	N.enthekiodon	Dorset
Ophthalmosaurus	Indeterminate	Dorset

Invertebrates

An aptychus with the name "Trigonellites latus", from the Kimmeridge Clay Formation

The invertebrate fauna of the Kimmeridge Clay includes:^[21]^[22]

Mollusca:
- Cardium striatulum
- Ostrea deltoidea
- Gryphaea (Exogyra) virgula
- The ammonite aptychus known as "Trigonellites latus"
- Belemnotheutis

Arthropoda:
- Phlyctosoma sp.
- Eryma sp.
- Magitalatimana sp.
- Mechochirus sp.
- Archaeolepas redtenbacheri

References

↑ Andy Lukas (17 July 2013). "Approaches to informed and balanced debate about shale gas in the UK – How we are working with the Communities" (PDF). Fracking and our Gas Future. AJ Lucas Group Ltd. p. 45. Retrieved 18 May 2016.
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Jurassic, Europe)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 545–549. ISBN 0-520-24209-2.
↑ Etches, S, Clarke, J. (2010). Life in Jurassic seas. Dorset, Dorchester: Epic Creative Print.
↑ http://www.theetchescollection.org/collections Retrieved 31/10/2017
1 2 Wilkinson, L.E.; Young, M.T.; Benton, M.J. (2008). "A new metriorhynchid crocodilian (Mesoeucrocodylia: Thalattosuchia) from the Kimmeridgian (Upper Jurassic) of Wiltshire, UK". Palaeontology. 51 (6): 1307–1333. doi:10.1111/j.1475-4983.2008.00818.x.
↑ Andrade, M.B.D.; Young, M.T.; Desojo, J.B.; Brusatte, S.L. (2010). "The evolution of extreme hypercarnivory in Metriorhynchidae (Mesoeucrocodylia: Thalattosuchia) based on evidence from microscopic denticle morphology". Journal of Vertebrate Paleontology. 30 (5): 1451–1465. doi:10.1080/02724634.2010.501442.
↑ Mark T. Young; Marco Brandalise De Andrade; Steve Etches; Brian L. Beatty (2013). "A new metriorhynchid crocodylomorph from the Lower Kimmeridge Clay Formation (Late Jurassic) of England, with implications for the evolution of dermatocranium ornamentation in Geosaurini". Zoological Journal of the Linnean Society. 169 (4): 820–848. doi:10.1111/zoj.12082.
↑ "Table 19.1," in Weishampel, et al. (2004). Page 415.
↑ Galton, Peter M. (1999). "Cranial anatomy of the hypsilophodont dinosaur Bugenasaura infernalis (Ornithischia: Ornithopoda) from the Upper Cretaceous of North America". Revue Paléobiologie, Genève. 18 (2): 517–534.
↑ "Table 13.1," in Weishampel, et al. (2004). Page 270.
1 2 Paul M. Barrett; Roger B.J. Benson; Paul Upchurch (2010). "Dinosaurs of Dorset: Part II, the sauropod dinosaurs (Saurischia, Sauropoda) with additional comments on the theropods". Proceedings of the Dorset Natural History and Archaeological Society. 131: 113–126.
↑ "Table 13.1," in Weishampel, et al. (2004). Page 267.
↑ "Table 13.1," in Weishampel, et al. (2004). Page 271.
↑ Benson, R.B.J. (2008). "New information on Stokesosaurus, a tyrannosauroid (Dinosauria: Theropoda) from North America and the United Kingdom". Journal of Vertebrate Paleontology. 28 (3): 732–750. doi:10.1671/0272-4634(2008)28[732:NIOSAT]2.0.CO;2.
↑ Brusatte, S.L.; Benson, R.B.J. (2013). "The systematics of Late Jurassic tyrannosauroids (Dinosauria: Theropoda) from Europe and North America". Acta Palaeontologica Polonica. 58 (1): 47–54. doi:10.4202/app.2011.0141.
↑ "A new monofenestratan pterosaur from the Kimmeridge Clay Formation (Kimmeridgian, Upper Jurassic) of Dorset, England - Acta Palaeontologica Polonica". www.app.pan.pl. Retrieved 2017-08-31.
↑ Benson, RBJ; Bowdler, T (2014). "Anatomy of Colymbosaurus (Reptilia, Plesiosauria) from the Kimmeridge Clay Formation of the U.K., and high diversity among Late Jurassic plesiosauroids". Journal of Vertebrate Paleontology. 34 (5): 1053–1071. doi:10.1080/02724634.2014.850087.
1 2 3 4 Roger B. J. Benson, Mark Evans, Adam S. Smith, Judyth Sassoon, Scott Moore-Faye, Hilary F. Ketchum and Richard Forrest (2013). "A Giant Pliosaurid Skull from the Late Jurassic of England". PLoS ONE. 8 (5): e65989. doi:10.1371/journal.pone.0065989. PMC 3669260. PMID 23741520.
1 2 3 Espen M. Knutsen (2012). "A taxonomic revision of the genus Pliosaurus (Owen, 1841a) Owen, 1841b". Norwegian Journal of Geology. 92 (2–3): 259–276. ISSN 0029-196X. Low resolution pdf Archived 2013-12-24 at the Wayback Machine. High resolution pdf Archived 2013-12-24 at the Wayback Machine.
↑ Noè, L. F.; Smith, D. T. J.; Walton, D. I. (2004). "A new species of Kimmeridgian pliosaur (Reptilia; Sauropterygia) and its bearing on the nomenclature of Liopleurodon macromerus". Proceedings of the Geologists' Association. 115: 13. doi:10.1016/S0016-7878(04)80031-2.
↑ http://www.fullbooks.com/The-Student-s-Elements-of-Geology7.html The Student's Elements of Geology by Sir Charles Lyell Part 7 out of 14 accessed 13 February 2009.
↑ Wignall, Paul B. (1990). "Benthic palaeoecology of the late Jurassic Kimmeridge Clay of England" (PDF). Special Papers in Palaeontology. The Palaeontological Association, London. 43. ISBN 978-0-901702-42-5. Retrieved February 8, 2011.

Bibliography

Galton, P.M. 1999. Cranial anatomy of the hypsilophodontid dinosaur Bugenasaura infernalis (Ornithischia: Ornithopoda) from the Upper Cretaceous of North America. Revue Pale´obiologie, 18, 517–534.