Enidae

Enidae
A live individual of Ena montana
Scientific classification
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
(unranked): clade Heterobranchia
clade Euthyneura
clade Panpulmonata
clade Eupulmonata
clade Stylommatophora
informal group Orthurethra
Superfamily: Enoidea
Family: Enidae
B. B. Woodward, 1903 (1880)
Genera

See text

Enidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks.

Anatomy

In this family, the number of haploid chromosomes lies between 21 and 25 (according to the values in this table).[1]

Taxonomy

1998 taxonomy

The following subfamilies of Enidae are based on work by Schileyko (1998)[2] and they are in use mainly by Russian malacologists:[3]

  • subfamily Pseudonapaeinae
  • subfamily Chondrulopsininae
  • subfamily Merdigerinae
  • subfamily Buliminusinae
  • subfamily Andronakiinae
  • subfamily Retowskiinae
  • subfamily Eninae - including Chondrula
  • subfamily Multidentulini

2005 taxonomy

The following two subfamilies have been recognized in the taxonomy of Bouchet & Rocroi (2005):

  • subfamily Eninae B. B. Woodward, 1903 (1880)
    • tribus Enini B. B. Woodward, 1903 (1880) - synonyms: Napaeinae A. J. Wagner, 1928; Jaminiinae Thiele, 1931; Pseudonapaeinae Schileyko, 1978; Retowskiinae Schileyko, 1978; Andronakiinae Schileyko, 1998
    • tribus Chondrulini Wenz, 1923
    • tribus Multidentulini Schileyko, 1978 - synonyms: Chondrulopsininae Schileyko, 1978; Merdigerinae Schileyko, 1984; Euchondrinae Schileyko, 1998
  • subfamily Buliminusinae Kobelt, 1880 - synonym Buliminidae Pfeiffer, 1879

Genera

Genera within the family Enidae include:

subfamilia Eninae

either in Enini or in Chondrulini

  • Brephulopsis Lindholm, 1925[3]
  • Caucasicola Hesse, 1917[3]
  • Chondrus Cuvier, 1817[3]
  • Georginapaeus Schileyko, 1988[3]
  • Peristoma Krynicki, 1833[3]
  • Ramusculus Lindholm, 1925[3]
  • Thoanteus Lindholm, 1925[3]
  • tribus Enini
    • Akramowskiella Schileyko, 1984[3]
    • Andronakia Lindholm, 1914[3]
    • Clausilioides Lindholm, 1925[3]
    • Ayna Páll-Gergely, 2009[4]
    • Differena Schileyko, 1984[3]
    • Ena Turton, 1831[3] - type genus of the family Enidae
    • Geminula Lindholm, 1925[3]
    • Imparietula Lindholm, 1925[3]
    • Jaminia Risso, 1826
    • Laevozebrinus Lindholm, 1925[3]
    • Ljudmilena Schileyko, 1984[3]
    • Mastoides Westerlund, 1896[3][5]
    • Megalena Hausdorf, 1999[6] There is only one species Megalena crassa (Retowski, 1887) in the genus Megalena.
    • Napaeus Albers, 1850
    • Ottorosenia Muratov, 1992[3]
    • Pseudochondrula Hesse, 1933[3]
    • Pseudonapaeus Westerlund, 1887[3]
    • Retowskia O. Boettger, 1881[3]
    • Subzebrinus Westerlund, 1887[3]
    • Turanena Lindholm, 1922[3]
  • tribus Chondrulini
  • tribus Multidentulini

subfamily Buliminusinae

  • Adzharia Hesse, 1933[3]
  • Buliminus Beck, 1837[3] - type genus of the subfamily Buliminusinae
  • Cyrenaeus Heller, 1971
  • Mordania Bank & Neubert, 1998
  • Paramastus Hesse, 1933
  • Pene Pallary, 1929

subfamily ?

  • Balearena Altaba, 2007[7] - type species † Balearena gymnesica Altaba, 2007[7]
  • Luchuena Habe, 1956

References

  1. Barker G. M.: Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. in Barker G. M. (ed.): The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, 2001, ISBN 0-85199-318-4. 1-146, cited pages: 139 and 142.
  2. Schileyko A. A. (1998). Treatise on Recent terrestrial pulmonate molluscs. Part 2. Gastrocoptidea, Hypselostomatidae, Vertiginidae, Truncatellinidae, Pachynodidae, Enidae, Sagdidae. Ruthenica Supplement 2(2): 129-261.
  3. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 Kantor Yu I., Vinarski M. V., Schileyko A. A. & Sysoev A. V.(published online on December 22, 2009). "Catalogue of the continental mollusks of Russia and adjacent territories". Version 2.3.
  4. Páll-Gergely, Barna (2009). "REVISION OF THE TURKISH RAMUSCULUS TAXA WITH DESCRIPTION OF AYNA GEN. NOV. (GASTROPODA: PULMONATA: ENIDAE)". 40 (1): 73–77.
  5. Schileyko A. A. (2007). "On the genus Mastoides Westerlund, 1896 (Pulmonata, Enidae)". Ruthenica 17(1-2): 85-87. abstract.
  6. Hausdorf H. (1999) "A new genus of the Buliminidae from Turkey (Gastropoda: S tylommatophora)" Journal of Natural History 33: 149-154. PDF Archived 2006-11-02 at the Wayback Machine.
  7. 1 2 Altaba C. R. (2007). "A new genus and species of Enidae (Gastopoda (sic!): Pulmonata) from the Quaternary of the Balearic Islands (Western Mediterranean)". Zootaxa 1595: 43-52. first page only.
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