Arachniodes

Arachniodes
Scientific classification
Kingdom:Plantae
Division:Pteridophyta
Class:Polypodiopsida/Pteridopsida
(disputed)
Order:Polypodiales
Family:Dryopteridaceae
Subfamily:Dryopteridoideae
Genus:Arachniodes
Blume 1828
Synonyms
  • Byrsopteris C.V.Morton 1960
  • Leptorumohra (H.Itô) H.Itô 1938
  • Polystichopsis (J.Sm.) Holttum 1947[1]

Arachniodes is a fern genus in the family Dryopteridaceae (wood ferns). A number of species in this genus are known as "holly ferns".[2]

The genus Arachniodes was first published by Carl Ludwig von Blume in 1828, with the single Indonesian species Arachniodes aspidioides.[3] But the genus wasn't widely recognized until Mary Douglas Tindale transferred the two species Byrsopteris amabilis and Byrsopteris aristata into it in 1961.[4]

Species

Arachniodes species include but are not limited to those in the following list. A species name preceded by (=) is considered by the referenced authority to be a synonym of the accepted species name listed above it.

1828 Blume

Carl Ludwig von Blume published a description of the first Arachniodes species in Enumeratio Plantarum Javae et Insularum Adjacentium (An Enumeration of the Plants of Java and Adjacent Islands) in 1828.

  • Arachniodes aspidioides Blume 1828[5]

1961 Tindale

The next Arachniodes species descriptions were published by Mary Douglas Tindale in Contributions from the New South Wales National Herbarium, volume 3, number 1, in 1961. This transferred two species from Conrad Vernon Morton's one-year-old genus Byrsopteris to Blume's 133-year-old genus Arachniodes.

Arachniodes aristata
(=) Byrsopteris amabilis (Blume) C.V.Morton 1960
(=) Polystichopsis amabilis (Blume) Tagawa 1958
(=) Rumohra amabilis (Blume) Ching 1934
(=) Nephrodium amabile (Blume) Hand.-Mazz. 1929
(=) Dryopteris amabilis (Blume) Kuntze 1891
(=) Polystichum amabile (Blume) J.Sm. 1866
(=) Aspidium amabile Blume 1828
Arachniodes aristata
  • Arachniodes aristata (G.Forst.) Tindale 1961 – East Indian holly fern[6]
    (India, Japan, South Korea, Sri Lanka)
(=) Byrsopteris aristata (G.Forst.) C.V.Morton 1960
(=) Polystichopsis aristata (G.Forst.) Tagawa 1959 (nom. illeg.)
(=) Polystichopsis aristata (G.Forst.) Holttum 1954
(=) Rumohra aristata (G.Forst.) Ching 1934
(=) Polystichum lobatum var. aristatum (G.Forst.) C.Chr. 1906
(=) Dryopteris aristata (G.Forst.) Kuntze 1891
(=) Lastrea aristata (G.Forst.) T. Moore 1858
(=) Polystichum aristatum (G.Forst.) C.Presl 1836
(=) Nephrodium aristatum (G.Forst.) C.Presl 1825
(=) Aspidium aristatum (G.Forst.) Sw. 1801
(=) Polypodium aristatum G.Forst. 1786

1962 Ohwi

Jisaburo Ohwi published the following Arachniodes species in the Journal of Japanese Botany, volume 37, number 3, in 1962.

  • Arachniodes assamica (Kuhn) Ohwi 1962[7] (Tibet, Yunnan, India, Nepal)
(=) Arachniodes assamica (Kuhn) Ching 1962 (nom. illeg.)
(=) Byrsopteris assamica (Kuhn) C.V. Morton 1960
(=) Polystichopsis assamica (Kuhn) Tagawa 1958
(=) Polystichum assamicum (Kuhn) Ching 1934
(=) Rumohra assamica (Kuhn) Ching 1934
(=) Dryopteris speciosa var. assamica (Kuhn) C.Chr. 1931
(=) Aspidium aristatum var. assamicum (Kuhn) Baker 1874
(=) Aspidium assamicum Kuhn 1869
Arachniodes simplicior
Arachniodes simplicior
  • Arachniodes cavalerii (H.Christ) Ohwi 1962[7]
(=) Arachniodes cavalerii (H.Christ) Ching 1962 (nom. illeg.)
(=) Polystichopsis cavalerii (H.Christ) Tagawa 1958
(=) Rumohra cavaleriei (H.Christ) Ching 1934
(=) Dryopteris cavalerii (H.Christ) C.Chr. 1905
(=) Aspidium cavalerii H.Christ 1904
  • Arachniodes maximowiczii (Baker) Ohwi 1962
(=) Arachniodes maximowiczii (Baker) Ching 1962 (nom. illeg.)
(=) Byrsopteris maximowiczii (Baker) C.V.Morton 1961
(=) Polystichopsis maximowiczii (Baker) Tagawa 1958
(=) Rumohra maximowiczii (Baker) Ching 1934
(=) Nephrodium maximowiczii Baker 1874
  • Arachniodes miqueliana (Maxim. ex Franch. & Sav.) Ohwi 1962
(=) Polystichopsis miqueliana (Maxim. ex Franch. & Sav.) Tagawa 1958
(=) Leptorumohra miqueliana (Maxim. ex Franch. & Sav.) H. Itô 1938[8]
(=) Rumohra miqueliana (Maxim. ex Franch. & Sav.) Ching 1934
(=) Dryopteris miqueliana (Maxim. ex Franch. & Sav.) C.Chr 1905
(=) Nephrodium miquelianum (Maxim. ex Franch. & Sav.) Kom. 1901
(=) Aspidium miquelianum Maxim. ex Franch. & Sav. 1876
  • Arachniodes mutica (Franch. & Sav.) Ohwi 1962
(=) Polystichopsis mutica (Franch. & Sav.) Tagawa 1958
(=) Leptorumohra mutica (Franch. & Sav.) Czerep. 1981[9]
(=) Rumohra mutica (Franch. & Sav.) Ching 1934
(=) Dryopteris mutica (Franch. & Sav.) C.Chr. 1905
(=) Aspidium muticum Franch. & Sav. 1876
Arachniodes standishii
Arachniodes standishii
  • Arachniodes nipponica (Rosenst.) Ohwi 1962
(=) Arachniodes nipponica (Rosenst.) Ching 1962 (nom. illeg.)
(=) Polystichopsis nipponica (Rosenst.) Tagawa 1958
(=) Rumohra nipponica (Rosenst.) Ching 1934
(=) Polystichum nipponicum Rosenst. 1914
  • Arachniodes pseudoaristata (Tagawa) Ohwi 1962
(=) Arachniodes pseudoaristata (Tagawa) Ching 1962
(=) Polystichopsis pseudoaristata (Tagawa) Tagawa 1958
(=) Rumohra pseudoaristata (Tagawa) H.Itô 1935
(=) Polystichum pseudoaristatum Tagawa 1932
  • Arachniodes simplicior (Makino) Ohwi 1962 – simpler holly fern[10]
(=) Arachniodes parasimplicior Ching ex Y.T.Hsieh 1991[11]
(=) Arachniodes simplicior (Makino) Ching 1962
(=) Polystichopsis simplicior (Makino) Tagawa 1958
(=) Rumohra simplicior (Makino) Ching 1934
(=) Polystichum simplicius (Makino) Tagawa 1932
(=) Polystichum aristatum var. simplicior (Makino) Matsum. 1904
(=) Aspidium aristatum var. simplicior Makino 1901
  • Arachniodes sinomiqueliana (Ching) Ohwi 1962
(=) Polystichopsis sinomiqueliana (Ching) Tagawa 1958
(=) Leptorumohra sinomiqueliana (Ching) Tagawa 1939[12]
(=) Rumohra sinomiqueliana Ching 1934
  • Arachniodes standishii (T.Moore) Ohwi 1962 – upside-down fern[13]
(=) Byrsopteris standishii (T. Moore) C.V. Morton 1961
(=) Polystichopsis standishii (T. Moore) Tagawa 1958
(=) Lastrea standishii T. Moore 1863

1962 Ching

The following species were transferred into Arachniodes by Ren Chang Ching in an article entitled "On the Nomenclature of the Compound-leaved Polysticha", published in the journal Acta Botanica Sinica, volume 10, number 3, Peking, China, September, 1962. Besides those listed below, a number of his species had already been published by Ohwi a few months earlier, and so appear in the section above, listed as synonyms marked nom. illeg.

  • Arachniodes amoena (Ching) Ching 1962 (Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Zhejiang)
(=) Arachniodes subamoena Ching 1986[14]
(=) Rumohra amoena Ching 1934
(=) Polystichum amabile var. controversum Hance ex Y.C.Wu 1932
  • Arachniodes arisanica (Rosenst.) Ching 1962
= Arachniodes globisora (Hayata) Ching 1964
  • Arachniodes carvifolia (Kunze) Ching 1962 – East Indian holly fern[15] (Yunnan, India, Philippines, Sri Lanka, U.S.)
(=) Byrsopteris carvifolia (Kunze) C.V.Morton 1960
(=) Rumohra carvifolia (Kunze) Ching 1934
(=) Dryopteris carvifolia (Kunze) C.Chr. 1924
(=) Polystichum carvifolium (Kunze) C.Chr. 1906 (nom. illeg.)
(=) Aspidium carvifolium Kunze 1848
  • Arachniodes chinensis (Rosenst.) Ching 1962
(=) Polystichopsis chinensis (Rosenst.) Holttum 1954
(=) Rumohra chinensis (Rosenst.) Ching 1934
(=) Polystichum amabile var. chinense Rosenst. 1914
  • Arachniodes coniifolia (T. Moore) Ching 1962 (Sichuan, Yunnan, Nepal)
(=) Byrsopteris coniifolia (T. Moore) C.V. Morton 1960
(=) Rumohra wallichii Ching 1934
(=) Lastrea coniifolia T. Moore 1857
(=) Aspidium coniifolium Wall. ex Kunze 1851 (nom. illeg.)
  • Arachniodes denticulata (Sw.) Ching 1962
(=) Arachniodes denticulata (Sw.) Proctor 1965 (nom. illeg.)
(=) Byrsopteris denticulata (Sw.) C.V.Morton 1960
(=) Lastrea denticulata (Sw.) T.Moore 1858
(=) Nephrodium denticulatum (Sw.) Fée 1852
(=) Rumohra denticulata (Sw.) Copel. 1947
(=) Dryopteris denticulata (Sw.) Kuntze 1891
(=) Nephrodium denticulatum (Sw.) Hook. 1862
(=) Polystichum denticulatum (Sw.) J.Sm. 1841
(=) Aspidium denticulatum (Sw.) Sw. 1801
(=) Polypodium denticulatum Sw. 1788
  • Arachniodes diffracta (Baker) Ching 1962
= Acrorumohra diffracta (Baker) H. Itô 1938 (Guangxi, Hainan, Yunnan, Japan, Vietnam)
(=) Polystichum diffractum (Baker) Masam. 1936
(=) Rumohra diffracta (Baker) Ching 1934
(=) Dryopteris diffracta (Baker) C. Chr. 1905
(=) Aspidium diffractum (Baker) H.Christ 1899
(=) Nephrodium diffractum Baker 1898
  • Arachniodes dimorphophyllum (Hayata) Ching 1962
(=) Rumohra dimorphophylla (Hayata) H. Itô 1935
(=) Polystichum dimorphophyllum Hayata 1911
  • Arachniodes exilis (Hance) Ching 1962
(=) Aspidium exile Hance 1883
  • Arachniodes festina (Hance) Ching 1962 (China)
(=) Arachniodes erythrosora Ching 1986
(=) Dryopteris festina (Hance) C.Chr. 1934
(=) Rumohra festina (Hance) Ching 1934
(=) Dryopteris fargesii (H.Christ) C. Chr. 1905
(=) Aspidium festinum Hance 1883
(=) Rumohra grossa Tardieu & C.Chr. 1938
  • Arachniodes hasseltii (Blume) Ching 1962
= Acrorumohra hasseltii (Blume) Ching 1964 (Hainan, Taiwan, India, Nepal)
(=) Byrsopteris hasseltii (Blume) C.V.Morton 1960
(=) Polystichopsis hasseltii (Blume) Holttum 1954
(=) Athyrium hasseltii (Blume) Ching 1940
(=) Rumohra hasseltii (Blume) Ching 1934
(=) Dryopteris hasseltii (Blume) C.Chr. 1905
(=) Phegopteris hasseltii (Blume) Mett. 1858
(=) Polypodium hasseltii Blume 1851
  • Arachniodes hainanensis (Ching) Ching 1962 (Hainan)
(=) Rumohra hainanensis Ching 1934
  • Arachniodes henryi (H.Christ) Ching 1962
(=) Byrsopteris henryi (H.Christ) C.V.Morton 1960
(=) Rumohra henryi (H.Christ) Ching 1934
(=) Dryopteris henryi (H.Christ) C.Chr. 1931
(=) Polystichum henryi H.Christ 1909
  • Arachniodes leucostegioides (C.Chr.) Ching 1962
(=) Byrsopteris leucostegioides (C.Chr.) C.V.Morton 1961
(=) Dryopteris leucostegioides C.Chr. 1920
  • Arachniodes nigrospinosa (Ching) Ching 1962
(=) Rumohra nigrospinosa (Ching) Ching 1934
(=) Polystichum nigrospinosum Ching 1931
  • Arachniodes obtusissima (Ching) Ching 1962
(=) Acrorumohra obtusissima (Mett. ex Kuhn) Ching 1964
(=) Rumohra obtusissima (Mett. ex Kuhn) Ching 1934
(=) Dryopteris obtusissima (Mett. ex Kuhn) H. Christ 1908
(=) Lastrea sparsa var. obtusissima (Mett. ex Kuhn) Bedd. 1876/1883
(=) Aspidium obtusissimum Mett. ex Kuhn 1869
  • Arachniodes simulans (Mett. ex Kuhn) Ching 1962
(=) Rumohra simulans Ching 1934
  • Arachniodes speciosa (D.Don) Ching 1962
(=) Byrsopteris speciosa (D.Don) C.V.Morton 1960
(=) Nephrodium speciousum (D.Don) Hand.-Mazz. 1929
(=) Dryopteris speciosa (D.Don) C. Chr. 1924
(=) Rumohra speciosa (D.Don) Ching 1934
(=) Polystichum speciosum (D.Don) J. Sm. 1842
(=) Aspidium speciosum D.Don 1825
  • Arachniodes spectabilis (Ching) Ching 1962
(=) Rumohra spectabilis Ching 1934
  • Arachniodes subreflexipinna (Ogata) Ching 1962
= Acrorumohra subreflexipinna (Ogata) H. Itô in Nakai & Honda 1938
(=) Rumohra subreflexipinna (Ogata) H.Itô 1935
(=) Dryopteris subreflexipinna Ogata 1935
  • Arachniodes tonkinensis (Ching) Ching 1962
(=) Arachniodes obtusipinnula Ching & Y.T. Xie 1984
(=) Rumohra tonkinensis Ching 1934
  • Arachniodes zeylanica (Ching) Ching 1962
(=) Rumohra zeylanica Ching 1934
(=) Aspidium undulatum Thwaites 1864 (nom. illeg.)

1962 Sa.Kurata

In 1962, Satoru Kurata published descriptions of the following species in the Science Report of the Yokosuka City Museum, Yokosuka, Japan.

  • Arachniodes × kenzo-satakei (Sa.Kurata) Sa.Kurata 1962
(=) Byrsopteris × kenzosatakei Sa.Kurata 1961

[16]

  • Arachniodes subamabilis Sa.Kurata 1962

1964 Ching & Chu H.Wang

Ren Chang Ching and Chu Hao Wang published the following species in Acta Phytotaxonomica Sinica, volume 9, number 4, 1964.

  • Arachniodes acuminata Ching & Chu H.Wang 1964
= Arachniodes sphaerosora (Tagawa) Ching 1965
  • Arachniodes caudata (Tagawa) Ching 1964
(=) Arachniodes australis Y.T.Hsieh 1991
(=) Polystichopsis simplicior var. major (Tagawa) Tagawa 1958
(=) Rumohra simplicior var. major (Tagawa) H. Itô 1935
(=) Polystichum simplicius var. majus Tagawa 1932
  • Arachniodes globisora (Hayata) Ching 1964
(=) Arachniodes arisanica (Rosenst.) Ching 1962
(=) Rumohra globisora (Hayata) H. Itô 1938
(=) Rumohra arisanica (Rosenst.) Ching 1934
(=) Polystichum arisanicum Rosenst. 1915
(=) Polystichum globisorum Hayata 1914
  • Arachniodes obtusiloba Ching & Chu H.Wang 1964
  • Arachniodes pseudocavalerii Ching & Chu H.Wang 1964
  • Arachniodes rhomboidea (Schott) Ching 1964
(=) Aspidium controversum Hance 1862
(=) Aspidium rhomboideum (Schott) Wall. ex Mett. 1858
(=) Polystichum rhomboideum Schott 1834
  • Arachniodes yoshinagae (Makino) Ching 1964
(=) Polystichum yoshinagae (Makino) Makino 1909
(=) Aspidium yoshinagae Makino 1899

1965 Ching

Ren Chang Ching in Acta Phytotaxonomica Sinica, volume 10, number 3, 1965, published:

  • Arachniodes sphaerosora (Tagawa) Ching 1965
(=) Arachniodes acuminata Ching & Chu H.Wang 1964[17]
(=) Dryopteris sphaerosora Tagawa 1934

1965 Sa.Kurata

Satoru Kurata published the following Arachniodes species name in the Journal of Geobotany (Hokuriku Journal of Botany), volume 13, in 1965.

  • Arachniodes hekiana Sa.Kurata 1965[18]

1965 Proctor

Published by George Richardson Proctor in the British Fern Gazette. volume 9, number 6, 1965

  • Arachniodes denticulata (Sw.) Proctor 1965 (nom. illeg.)
= Arachniodes denticulata (Sw.) Ching 1962
  • Arachniodes rigidissima (Hook.) Proctor 1965
(=) Byrsopteris rigidissima (Hook.) C.V. Morton 1960
(=) Dryopteris rigidissima (Hook.) C. Chr. 1920
(=) Polystichum denticulatum var. rigidissima (Hook.) Hieron. 1904
(=) Nephrodium denticulatum var. rigidissimum Hook. 1860

1967 Schelpe

Edmund André Charles Louis Eloi Schelpe published the following species in Boletim da Sociedade Broteriana, sér. 2, vol. 41, 1967.

  • Arachniodes webbiana Schelpe 1967
(=) Aspidium webbianum A. Braun 1841
  • Arachniodes foliosa Schelpe 1967
(=) Arachniodes webbiana var. foliosa (C. Chr.) Gibby, Rasbach, Reichst., Widén & Viane 1992
(=) Dryopteris foliosa C. Chr. 1937

1968 Sa.Kurata

In 1968, Satoru Kurata published the following Arachniodes species names in the Journal of Geobotany (Hokuriku Journal of Botany), volume 16.

  • Arachniodes repens Sa.Kurata 1968[19]
  • Arachniodes yasu-inouei Sa.Kurata 1968
  • Arachniodes × masakii Sa.Kurata 1968
  • Arachniodes × clivorum Sa.Kurata 1968
  • Arachniodes × kurosawae Shimura & Sa.Kurata in Sa.Kurata 1968
  • Arachniodes × pseudohekiana Sa.Kurata 1968
  • Arachniodes × sasamotoi Sa.Kurata 1968

1969 & 1970 Nakaike

Toshiyuki Nakaike published the following new species in Journal of Geobotany (Hokuriku Journal of Botany), volume 17, 1969, and volume 18, 1970.

  • Arachniodes okinawensis Nakaike 1969 (Japan)
  • Arachniodes pseudorepens Nakaike 1970 (Japan)

1974 Ching

In Flora Tsinlingensis, volume 2, 1974, Ren Chang Ching published:

  • Arachniodes centrochinensis Ching 1974
= Arachniodes simulans (Ching) Ching 1962 (China)

1975 R.M.Tryon & D.S.Conant

Rolla Milton Tryon and Davis S. Conant published the following species in Acta Amazonica, volume 5, number 1, in 1975.

  • Arachniodes macrostegia (Hook.) R.M.Tryon & D.S.Conant 1775 (Brazil, Venezuela)
(=) Arachniodes macrostegia (Hook.) Proctor 1982 (nom. illeg.)
(=) Dryopteris macrostegia (Hook.) Kuntze 1891
(=) Nephrodium macrostegium Hook. 1862
Arachniodes sporadosora

1975 Nakaike

In 1975, Toshiyuki Nakaike published the following species names in his Enumeratio Pteridophytarum Japonicarum, Filicales.

  • Arachniodes sporadosora (Kunze) Nakaike 1775
(=) Aspidium sporadosorum Kunze 1848
  • Arachniodes japonica (Kurata) Nakaike 1775
(=) Arachniodes chinensis var. japonica Sa.Kurata 1965

1978 Sa.Kurata

Satoru Kurata published the following Arachniodes species names in the Journal of Geobotany (Hokuriku Journal of Botany), volume 25, in 1978.

  • Arachniodes cantilenae Sa.Kurata 1978
  • Arachniodes hiugana Sa.Kurata 1978
  • Arachniodes kenzo-satakei Sa.Kurata 1978
  • Arachniodes tomitae Sa.Kurata 1978
  • Arachniodes × minamitanii Sa.Kurata 1978 (A. hiugana × A. nipponica)
  • Arachniodes × mirabilis Sa.Kurata 1978 (A. assamica × A. rhomboidea)
  • Arachniodes × mitsuyoshiana Sa.Kurata 1978 (A. miqueliana × A. simplicior)
  • Arachniodes × respiciens Sa.Kurata 1978 (A. exilis × A. simplicior)

1982 Proctor

  • Arachniodes argillicola (Proctor) Proctor 1982
(=) Polystichopsis argillicola Proctor 1961
  • Arachniodes chaerophylloides (Poir.) Proctor 1982 – false holly fern[20]
(=) Polystichopsis chaerophylloides (Poir.) C.V.Morton 1960
(=) Lastreopsis chaerophylloides (Poir.) Tindale 1957
(=) Thelypteris chaerophylloides (Poir.) Proctor 1953
(=) Rumohra chaerophylloides (Poir.) Ching 1934
(=) Dryopteris chaerophylloides (Poir.) C.Chr. 1920
(=) Polypodium chaerophylloides Poir. 1804

1983 Y.T.Hsieh

Yin Tang Hsieh in Acta Botanica Yunnanica, volume 5, number 1, 1983.

  • Arachniodes jinpingensis Y.T.Hsieh 1983 (Yunnan)
(=) Arachniodes chinensis var. jinpingensis (Y.T. Hsieh) S.K.Wu & H.He 1996
  • Arachniodes valida Y.T.Hsieh 1983 (Yunnan)
  • Arachniodes dayaoensis Y.T.Hsieh 1983 (Yunnan)

1984 Y.T.Hsieh, Ching, et al.

Yin Tang Hsieh in Bulletin of Botanical Research, volume 4, number 2, 1984.

  • Arachniodes caudifolia Ching & Y.T.Hsieh 1984 (Yunnan)
  • Arachniodes gansuensis (Ching) Y.T.Hsieh 1984 (Yunnan, Sichuan)
(=) Arachniodes caudata var. kansuensis Ching 1974
  • Arachniodes guangnanensis Y.T.Hsieh 1984 (Yunnan)
  • Arachniodes lushuiensis Y.T.Hsieh 1984 (Yunnan)
  • Arachniodes reducta Y.T.Hsieh & Y.P.Wu 1984 (Zhejiang)
  • Arachniodes subaristata Ching & Y.T.Hsieh 1984 (China)

Yin Tang Hsieh and Ren Chang Ching, in Acta Phytotaxonomica Sinica, volume 22, number 2, 1984.

  • Arachniodes ishingensis Ching & Y.T.Hsieh 1984
(=) Arachniodes yixinensis Ching & Y.T.Hsieh 1984
  • Arachniodes obtusipinnula Ching & Y.T.Xie 1984
= Arachniodes tonkinensis (Ching) Ching 1962
  • Arachniodes ziyunshanensis Y.T.Hsieh 1984

Ren Chang Ching and Zheng-yu Liu, in Acta Phytotaxonomica Sinica, volume 22, number 4, 1984.

  • Arachniodes nanchuanensis Ching & Z.Y.Liu 1984 (Sichuan)

1986 Ching

Ren Chang Ching published the following species names in the Bulletin of Botanical Research, volume 6, number 3, in 1986.

  • Arachniodes abrupta Ching 1986 (Guangxi, Zhejiang)
  • Arachniodes ailaoshanensis Ching 1986 (Yunnan)
  • Arachniodes anshunensis Ching & Y.T. Hsieh 1986 (Guizhou)
  • Arachniodes aristatissima Ching 1986 (Zhejiang)
  • Arachniodes attenuata Ching 1986 (Yunnan)
  • Arachniodes austro-yunnanensis Ching 1986 (Yunnan)
  • Arachniodes baiseensis Ching 1986 (Guangxi)
  • Arachniodes calcarata Ching 1986 (Guizhou, Hunan, Sichuan)
  • Arachniodes cyrtomifolia Ching 1986 (Sichuan)
  • Arachniodes cornopteris Ching 1986 (Sichuan)
  • Arachniodes costulisora Ching 1986 (Sichuan)
  • Arachniodes decomposita Ching 1986 (Guizhou)
  • Arachniodes elevatas Ching 1986 (Sichuan)
  • Arachniodes emeiensis Ching 1986 (Sichuan)
  • Arachniodes erythrosora Ching 1986
= Arachniodes festina (Hance) Ching 1962 (Guangdong)
  • Arachniodes falcata Ching 1986 (Sichuan)
  • Arachniodes fengii Ching 1986
  • Arachniodes foeniculacea Ching 1986
  • Arachniodes fujianensis Ching 1986
  • Arachniodes gigantea Ching 1986
  • Arachniodes gijiangensis Ching 1986
  • Arachniodes gizushanensis Ching 1986
  • Arachniodes gongshanensis Ching & Y.T.Hsieh 1986
  • Arachniodes gradata Ching 1986
  • Arachniodes guangtongensis Ching 1986
  • Arachniodes guangxiensis Ching 1986
  • Arachniodes guanxianensis Ching 1986
  • Arachniodes hekouensis Ching 1986
  • Arachniodes heyuanensis Ching 1986
  • Arachniodes huapingensis Ching 1986
  • Arachniodes hunanensis Ching 1986
  • Arachniodes jiangxiensis Ching 1986
  • Arachniodes jinfoshanensis Ching 1986
  • Arachniodes jingdongensis Ching 1986
  • Arachniodes leuconeura Ching 1986
  • Arachniodes liyangensis Ching 1986
  • Arachniodes longipinna Ching 1986
  • Arachniodes lushanensis Ching 1986
  • Arachniodes maoshanensis Ching 1986
  • Arachniodes mengziensis Ching 1986
  • Arachniodes multifida Ching 1986
  • Arachniodes nanqingensis Ching 1986
  • Arachniodes neoaristata Ching 1986
  • Arachniodes nitidula Ching 1986
  • Arachniodes pianmaensis Ching 1986
  • Arachniodes pseudo-assamica Ching 1986
  • Arachniodes pseudo-longipinna Ching 1986
  • Arachniodes pseudo-simplicior Ching 1986
  • Arachniodes semifertilis Ching 1986
  • Arachniodes setifera Ching 1986
  • Arachniodes sichuanensis Ching 1986
  • Arachniodes similis Ching 1986
  • Arachniodes sino-aristata Ching 1986
  • Arachniodes sino-rhomboidea Ching 1986
  • Arachniodes shuangbaiensis Ching 1986
  • Arachniodes sparsa Ching 1986
  • Arachniodes spino-serrulata Ching 1986
  • Arachniodes subamoena Ching 1986
= Arachniodes amoena (Ching) Ching 1962
  • Arachniodes triangularis Ching 1986
  • Arachniodes xinpingensis Ching 1986
  • Arachniodes yaomashanensis Ching 1986
  • Arachniodes yinjiangensis Ching 1986
  • Arachniodes yunnanensis Ching 1986

1986 Seriz.

  • Arachniodes borealis Seriz. 1986 (Japan, Korea)[21]
= Leptorumohra borealis ( Seriz. ) Shmakov 2009

1986 Y.T.Hsieh

Yin Tang Hsieh published the following species names in the Bulletin of Botanical Research, volume 6, number 4, in 1986.

  • Arachniodes chingii Y.T.Hsieh 1986
  • Arachniodes damiaoshanensis Y.T.Hsieh 1986
  • Arachniodes futeshanensis Y.T.Hsieh 1986
  • Arachniodes nibashanensis Y.T.Hsieh 1986
  • Arachniodes yunqiensis Y.T.Hsieh 1986

1991 Ching & Hsieh

Ren Chan Ching and Yin Tang Hsieh, in the Bulletin of Botanical Research volume 11, number 2, 1991, published:

  • Arachniodes basipinnata (Ching) Ching ex Y.T.Hsieh 1991
(=) Rumohra assamica fo. basipinnata Ching 1934
(=) Arachniodes assamica fo. basipinnata Ching 1934 (nom. illeg.)
  • Arachniodes lanceolata Y.T.Hsieh 1991
  • Arachniodes menglianensis Y.T.Hsieh 1991
  • Arachniodes parasimplicior Ching ex Y.T.Hsieh 1991
= Arachniodes simplicior (Makino) Ohwi 1962
  • Arachniodes fengyangshanensis Ching & C.F.Zhang ex Y.T.Hsieh 1991
= Arachniodes exilis (Hance) Ching 1962

1992 Nakaike

  • Arachniodes davalliaeformis (Christ) Nakaike 1992[22] – shiny bristle fern
(=) Aspidium aristatum var. davalliaeforme (H.Christ) H.Christ

[23]

1993 W.H.Wagner

  • Arachniodes insularis W.H.Wagner 1993 – Santa Cruz Island holly fern[24]

1995 S.F.Wu

The following two species were published by Shi Fu Wu in Keys to the Vascular Plants of the Wuling Mountains Area (edited by Wang Wentsai) 572–573, pl. 5. Beijing 1995.

  • Arachniodes hupingshanensis S.F.Wu 1995 (China)
  • Arachniodes wulingshanensis S.F.Wu 1995 (China)

1998 R.C.Moran & B.Øllg.

2008 Fraser-Jenk.

In 2008, Christopher Roy Fraser-Jenkins published the following new Arachniodes species in his Taxonomic revision of three hundred Indian subcontinental pteridophytes with a revised census-list: a new picture of fern-taxonomy and nomenclature in the Indian subcontinent.

  • Arachniodes cornucervi (D.Don) Fraser-Jenk. 2008
(=) Aspidium cornucervi D.Don 1825
  • Arachniodes palmipes (Kunze) Fraser-Jenk. 2008
(=) Aspidium palmipes Kunze 1851
  • Arachniodes sledgei Fraser-Jenk. 2008

Leptorumohra

The genus Leptorumohra was segregated from the genus Rumohra in 1938 by Hiroshi Itô.[25] In 1962, Jisaburo Ohwi included at least three of the Leptorumohra species in Arachniodes, but these placements were not universally accepted.[25] In 2006, Alan Reid Smith & al. in "A classification for extant ferns" included the genus Leptorumohra in Arachniodes.[26] Leptorumohra species, which may be considered part of the genus Arachniodes, include:

(=) Dryopteris amurensis H.Christ 1909
  • Arachniodes borealis Seriz. 1986
(=) Leptorumohra borealis ( Seriz.) Shmakov 2009[27]
  • Arachniodes miqueliana (Maxim. ex Franch. & Sav.) Ohwi 1962
(=) Polystichopsis miqueliana (Maxim. ex Franch. & Sav.) Tagawa 1958
(=) Leptorumohra miqueliana (Maxim. ex Franch. & Sav.) H. Itô 1938[8]
(=) Rumohra miqueliana (Maxim. ex Franch. & Sav.) Ching 1934
(=) Dryopteris miqueliana (Maxim. ex Franch. & Sav.) C.Chr 1905
(=) Nephrodium miquelianum (Maxim. ex Franch. & Sav.) Kom. 1901
(=) Aspidium miquelianum Maxim. ex Franch. & Sav. 1876
  • Arachniodes mutica (Franch. & Sav.) Ohwi 1962
(=) Polystichopsis mutica (Franch. & Sav.) Tagawa 1958
(=) Leptorumohra mutica (Franch. & Sav.) Czerep. 1981[9]
(=) Rumohra mutica (Franch. & Sav.) Ching 1934
(=) Dryopteris mutica (Franch. & Sav.) C.Chr. 1905
(=) Aspidium muticum Franch. & Sav. 1876
  • Arachniodes quadripinnata (Hayata) Seriz. 1986[28]
(=) Leptorumohra quadripinnata (Hayata) H.Itô 1938[29]
(=) Microlepia quadripinnata Hayata 1911
  • Arachniodes sinomiqueliana (Ching) Ohwi 1962
(=) Polystichopsis sinomiqueliana (Ching) Tagawa 1958
(=) Leptorumohra sinomiqueliana (Ching) Tagawa 1939[12]
(=) Rumohra sinomiqueliana Ching 1934

References

  1. ↑ Arachniodes Tropicos.org. Missouri Botanical Garden. 26 Jan 2012
  2. ↑ Arachniodes Flora of North America 26 Jan 2012
  3. ↑ Blume, Carl (Karl) Ludwig von; Enumeratio Plantarum Javae fasc. 2: 241. 1828.
  4. ↑ Tindale, Mary Douglas; Contributions from the New South Wales National Herbarium 3(1): 90. 1961.
  5. ↑ Blume, Carl (Karl) Ludwig von: Enumeratio Plantarum Javae fasc. 2: 241. 1828.
  6. ↑ A. aristata Archived April 3, 2012, at the Wayback Machine. Hardy fern library 26 Jan 2012
  7. 1 2 Ohwi, Jisaburo: Journal of Japanese Botany 37(3): 76. 1962.
  8. 1 2 A. miqueliana Tropicos.org. Missouri Botanical Garden. 26 Jan 2012
  9. 1 2 A. mutica Tropicos.org. Missouri Botanical Garden. 26 Jan 2012
  10. ↑ "Arachniodes simplicior". Natural Resources Conservation Service PLANTS Database. USDA. Retrieved 26 Jan 2012.
  11. ↑ A. parasimplicior Tropicos.org. Missouri Botanical Garden. 26 Jan 2012
  12. 1 2 A. sinomiqueliana Tropicos.org. Missouri Botanical Garden. 26 Jan 2012
  13. ↑ A. standishii Archived April 15, 2012, at the Wayback Machine. Hardy fern library 26 Jan 2012
  14. ↑ A. subamoena Tropicos.org. Missouri Botanical Garden. 26 Jan 2012
  15. ↑ A. carvifolia Ensyclopedia of Life (EOL) 27 Jan 2012
  16. ↑ Kurata, Satoru: Journal of Geobotany/Hokuriku Journal of Botany 10: 36. 1961.
  17. ↑ A. acuminata Tropicos.org. Missouri Botanical Garden. 26 Jan 2012
  18. ↑ Kurata, Satoru: Journal of Geobotany/Hokuriku Journal of Botany 13: 99–100. 1965.
  19. ↑ Kurata, Satoru: Journal of Geobotany/Hokuriku Journal of Botany 16: 98. 1968.
  20. ↑ "Arachniodes chaerophylloides". Natural Resources Conservation Service PLANTS Database. USDA. Retrieved 26 Jan 2012.
  21. ↑ A. borealis International Plant Names Index (IPNI) 30 Jan 2012
  22. ↑ Arachniodes World species list. 27 Jan 2012
  23. ↑ A. davalliaeformis International Plant Names Index (IPNI) 27 Jan 2012
  24. ↑ "Arachniodes insularis". Natural Resources Conservation Service PLANTS Database. USDA. Retrieved 26 Jan 2012.
  25. 1 2 Leptorumohra Tropicos.org. Missouri Botanical Garden. 26 Jan 2012
  26. ↑ Smith & al. Smith, A.R.; Pryer, K.M.; Schuettpelz, E.; Korall, P.; Schneider, H.; Wolf, P.G. "A classification for extant ferns". Taxon 55: 705–731. (2006)
  27. ↑ Leptorumohra borealis International Plant Names Index (IPNI) 26 Dec 2011
  28. ↑ A. quadripinnata International Plant Names Index (IPNI) 26 Jan 2012
  29. ↑ Leptorumohra quadripinnata Tropicos.org. Missouri Botanical Garden. 26 Jan 2012
This article is issued from Wikipedia. The text is licensed under Creative Commons - Attribution - Sharealike. Additional terms may apply for the media files.