Sea eagle

Sea eagles vary in size, from Sanford's sea eagle, averaging 2.0–2.7 kg (4 lb 7 oz–5 lb 15 oz), to the huge Steller's sea eagle, weighing up to 9 kg (20 lb).[2] At up to 6.9 kg (15 lb 3 oz), the white-tailed eagle is the largest eagle in Europe. Bald eagles can weigh up to 6.3 kg (13 lb 14 oz), making them the largest eagle native to North America. There are exceptional records of even heavier individuals in both the white-tailed and bald eagles, although not surpassing the largest Steller's sea eagles. The white-bellied sea eagle can weigh up to 4.5 kg (9 lb 15 oz).[2] They are generally overall brown (from rich brown to dull grey-brown), often with white to the head, tail or underparts. Some of the species have an all-yellow beak as adults, which is unusual among eagles.[2]

Their diets consist mainly of fish, aquatic birds and small mammals. Nests are typically very large and positioned in a tree, but sometimes on a cliff.[2]

The genus Haliaeetus was introduced in 1809 by the French naturalist Marie Jules César Savigny in his chapter on birds in the Description de l'Égypte.[3][4] The two fish eagles in the genus Ichthyophaga found to lie within the genus in a genetic study in 2005,[5] and then placed therein. They are very similar to the tropical Haliaeetus species. A prehistoric (i.e. extinct before 1500) form from Maui in the Hawaiian Islands may represent a species or subspecies in this genus.

The 10 living species are:[2]

Image	Scientific name	Common Name	Distribution
	Haliaeetus leucogaster	White-bellied sea eagle	India and Sri Lanka through Southeast Asia to Australia
	Haliaeetus sanfordi	Sanford's sea eagle	Solomon Islands
	Haliaeetus vocifer	African fish eagle	sub-Saharan Africa
	Haliaeetus vociferoides	Madagascan fish eagle	Madagascar
	Haliaeetus leucoryphus	Pallas's fish eagle	Central Asia, between the Caspian Sea and the Yellow Sea, from Kazakhstan and Mongolia to the Himalayas, Bangladesh and northern India.
	Haliaeetus albicilla	White-tailed eagle	Western Greenland, most of Eurasia as far south as southeastern China and Japan
	Haliaeetus leucocephalus	Bald eagle	Canada, all of the continental United States, and northern Mexico
	Haliaeetus pelagicus	Steller's sea eagle	coastal northeastern Asia
	Haliaeetus humilis	Lesser fish eagle	Kashmir through southeast India, Nepal, and Burma towards Indochina
	Haliaeetus ichthyaetus	Grey-headed fish eagle	South East Asia

The tail is entirely white in adult Haliaeetus species except for Sanford's, White-bellied, and Pallas's. Three species pairs exist: white-tailed and bald eagles, Sanford's and white-bellied sea eagles, and the African and Madagascan fish eagles,[6] each of these consists of a white- and a tan-headed species.

Haliaeetus is possibly one of the oldest genera of living birds. A distal left tarsometatarsus (DPC 1652) recovered from early Oligocene deposits of Fayyum, Egypt (Jebel Qatrani Formation, about 33 million years ago (Mya) is similar in general pattern and some details to that of a modern sea eagle.[7] The genus was present in the middle Miocene (12-16 Mya) with certainty.[8]

The relationships to other genera in the family are less clear; they have long been considered closer to the genus Milvus (kites) than to the true eagles in the genus Aquila on the basis of their morphology and display behaviour,[2][9] more recent genetic evidence agrees with this, but points to them being related to the genus Buteo (buzzards/hawks), as well, a relationship not previously thought close.[6]

The origin of the sea eagles and fishing eagles is probably in the general area of the Bay of Bengal. During the Eocene/Oligocene, as the Indian subcontinent slowly collided with Eurasia, this was a vast expanse of fairly shallow ocean; the initial sea eagle divergence seems to have resulted in the four tropical (and Southern Hemisphere subtropical) species living around the Indian Ocean today. The Central Asian Pallas's sea eagle's relationships to the other taxa is more obscure; it seems closer to the three Holarctic species which evolved later and may be an early offshoot of this northward expansion; it does not have the hefty yellow bill of the northern forms, retaining a smaller, darker beak like the tropical species.[6]

The rate of molecular evolution in Haliaeetus is fairly slow, as is to be expected in long-lived birds which take years to successfully reproduce. In the mtDNA cytochrome b gene, a mutation rate of 0.5–0.7% per million years (if assuming an Early Miocene divergence) or maybe as little as 0.25–0.3% per million years (for a Late Eocene divergence) has been shown.[6]

A 2005 molecular study found that the genus is paraphyletic and subsumes Ichthyophaga, the species diverging into a temperate and tropical group.[10]

Nesting pairs of both the bald eagle and white-bellied sea eagle have been subject to live-streaming webcam footage.[11][12]

• Brahminy kite, also called red-backed sea eagle

Sources

• del Hoyo, J.; Elliott, A.; Sargatal, J., eds. (1994). Handbook of the Birds of the World. 2. Barcelona: Lynx Edicions. ISBN 84-87334-15-6.CS1 maint: ref=harv (link)
• Wink, M.; Heidrich, P.; Fentzloff, C. (1996). "A mtDNA phylogeny of sea eagles (genus Haliaeetus) based on nucleotide sequences of the cytochrome b gene" (PDF). Biochemical Systematics and Ecology. 24 (7–8): 783–791. doi:10.1016/S0305-1978(96)00049-X.CS1 maint: ref=harv (link)