Paleobiota of the Posidonia Shale

The Posidonia Shale or Posidonienschiefer Formation is a geological formation of southwestern Germany, northern Switzerland, western Czech Republic, northwestern Austria, southeast Luxembourg and the Netherlands, that spans about 3 million years during the Early Jurassic period (early Toarcian stage). It is known for its detailed fossils, especially sea fauna, listed below.[1] Composed mostly by black shale, the formation is a Lagerstätte, where fossils show exceptional preservation (Including exquisite soft tissues), with a thickness that varies from about 1 m to about 40 m on the Rhine level, being on the main quarry at Holzmaden between 5 and 14 m.[1] Some of the preserved material has been transformed into fossil hydrocarbon Jet, specially wood remains, used for jewelry.[2] The exceptional preservation seen on the Posidonia Shale has been studied since the late 1800s, finding that a cocktail of chemical and environmental factors let to such an impressive conservation of the marine fauna.[2] The most common theory is the changes on the oxygen level, where the different anoxic events of the Toarcian left oxygen-depleted bottom waters, with the biota dying and falling to the bottom without any predator able to eat the dead bodies.[3]

18x6 m fossilized floating wood (Araucarioxylon), with Crinoids attached (Pentacrinites & Seirocrinus). It is one of the most emblematic fossils of the formation, where the anoxic seas of the lower toarcian lead to an exquisite preservation.

Invertebrata

Ichnofossils

Genus	Species	Location	Material	Notes	Images
Thalassinoides[52]	Thalassinoides sp.	Dotternhausen Holzmaden Ohmden Altdorf Banz Hemmikon Maurach	Burrowing and track ichnofossils	Burrow-like ichnofossils, that can be related to Crustaceans, Annelids and Fishes.[52] The presence of this burrows changed along the different depositional layers, interpreted as result of relative magnitudes and durations of a series oxygenation events.[52] Increased Oxygen conditions eventually led to a level that permitted both the survival of larger Chondrites and Thalassinoides producing organisms, as well the depth of the Burrow-like structures.[52] The changes on the layers are detailed enough to know that oxygenation-change events duration was sufficient to allowe the migration and establishment of trace-producing organisms, establishing an "equilibrium" with bottom-water oxygen conditions.[52]	Thalassinoides found on the Posidonia Shale.
Chondrites[53]	Chondrites bollensis	Banz Aichelberg Dotternhausen Holzmaden Ohmden Maurach Hemmikon Dudelange-Zoufftgen	Burrowing and track ichnofossils	Burrow-like ichnofossils. Interpreted as the feeding burrow of a sediment-ingesting animal.[54] A more recent study has find that Scoloplos armiger and Heteromastus filiformis, occurring in the German Wadden Sea in the lower parts of tidal flats, make burrows that are homonymous with numerous trace fossils of the ichnogenus.[55]	Illustration of Chondrites bollensis
Phymatoderma[56]	Phymatoderma granulata	Banz Altdorf Dotternhausen Holzmaden Ohmden Maurach Hemmikon	Burrowing and track ichnofossils.	Burrow-like ichnofossils. It consists commonly on a subhorizontal branching burrow system consisting of radiating tunnels filled with fecal pellets.[56] It has been interpreted as a product made by an Endobenthic deposit-feeding animal, specially a Fodinichnia, burrows produced by benthonic subsurface food-mining activity, as is proved by the tunnels and pelletal infill.[56] The study of the Fecal Pellets has revelated that the maker of this ichnogenus was an epicontinental shelf setting non-selective deposit feeder, ingesting particles on the sediment surface without selection. A mode of feeding common on aquatic Benthos, reported on modern animals such as Spionid Polychaete worms, tropical Holothurians and Spatangoid urchins.[57]
Zoophycos[56][58]	Zoophycos sp.	Banz Dotternhausen Holzmaden Ohmden Maurach Hemmikon	Burrowing and track ichnofossils.[59]	Burrow-like ichnofossils. It has been related to Echiuran annelids,[60] but also from moving and feeding polychaete worms.[61]	Example of Zoophycos fossil
Fucoides[62]	Fucoides bollensis	Banz Dotternhausen Holzmaden Ohmden Maurach Hemmikon	Burrowing and track ichnofossils.	Burrow-like ichnofossils. Considered an "Algae incertae sedis" fossil, it was recovered subdivided into 16 different groups, being in 1880 along certain ichnofossil genera, such as Cruziana with a massive nomenclatural complexity. Considered now a feeding burrow of a sediment-ingesting animal, sometimes synonymized with Chondrites.
Planolites[63]	Planolites montanus Planolites beverleyensis	Banz Dotternhausen Holzmaden Ohmden Maurach Hemmikon	Burrowing and track ichnofossils.	Burrow-like ichnofossils. It is controversial, since is considered a strictly a junior synonym of Palaeophycus.[64]	Example of Planolites fossil
Palaeophycus[65]	Palaeophycus tubularis	Dotterhausen Irlbach	Burrowing and track ichnofossils.	Burrow-like ichnofossils. Palaeophycus is considered related with Planolites, being a litoral fodichnia, probably from a Priapulidan.	Example of Palaeophycus fossil
Helminthopsis[65]	Helminthopsis isp.	Irlbach	Burrowing and track ichnofossils.	Burrow-like ichnofossils. It is interpreted as a grazing trail or Fodinichnia, produced at shallow depth in sediment by Polychaetes and Priapulids.[65]	Example of Helminthopsis fossil
Gyrochorte[65]	Gyrochorte isp.	Irlbach	Burrowing and track ichnofossils.	Burrow-like ichnofossils. Gyrochorte is interpreted as a result of active digging on the sediment by deposit-feeding worm-like animal, probably an Annelid or similar kinds of creatures, such as Crustaceans, Sea Urchins, nearshore fishes, etc.
Cylindrichnus[65]	Cylindrichnus isp.	Altdorf Irlbach	Burrowing and track ichnofossils.	Burrow-like ichnofossils. Cylindrichnus isp. was found only on seashore-related sections, and probably represents litoral Polychaete Burrows.[66]
Asterosoma[65]	Asterosoma ludwigae	Holzmaden Ohmden	Burrowing and track ichnofossils.	Burrow-like ichnofossils. Vertical or oblique complex trace fossil composed of a bunch of spindle-shaped structures and associated tubes, typical of a restricted environment (?estuarine/lagoonal).
Spongeliomorpha[65]	Spongeliomorpha isp.	Holzmaden Dotterhausen Irlbach	Burrows and associated traces	Burrow-like ichnofossils. Spongeliomorpha is believed to come from the domicile of Crustaceans: Anomuras (Probably Eocarcinoidea) and Decapodans (Probably Glypheidae), created as they dig in a firm, semiconsolidated substrate.
Diplocraterion[65]	Diplocraterion parallelum	Holzmaden Ohmden Altdorf Dotternhausen Irlbach	Burrows and associated traces	Burrow-like ichnofossils. Most Diplocraterion show only protrusive spreit, like the local ones, produced under predominantly erosive conditions where the organism was constantly burrowing deeper into the substrate as sediment was eroded from the top. It can be Made by Crustaceans, Annelids or other benthic fauna.[65]	Diplocraterion parallelum diagram

Anthozoa

Genus	Species	Location	Material	Notes	Images
Thecocyathus[27][67]	Thecocyathus mitrae Thecocyathus tintinabulum Thecocyathus mactra	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Irlbach Maurach Hemmikon Dudelange-Zoufftgen	Polyps	A stone cora of the family Caryophylliidae inside Hexacorallia. Related to shallow waters, this genus is the main coral found on the Posidonia Shale, resembling the modern Polycyathus muellerae. Its fossils are related with near-land facies, Coralline Islands and relatively small landmases shuch as the Bohemian Massif.	Thecocyathus mitrae from Banz

Porifera

Genus	Species	Location	Material	Notes	Images
Cribrospongia[27]	Cribrospongia (Tremadictyon) rugatum	Banz Altdorf Mistelgau Aichelberg Schandelah Dotternhausen Hemmikon	Specimens	A sea sponge of the family Cribrospongiidae inside Sceptrulophora. Found on Shallow and basinal waters, some specimens get 22 cm wide, with a funnel-like morphology. It is relatively common on nearshore strata, but generally rare.	Cribrospongia specimen
Stauroderma[27]	Stauroderma lochensis Stauroderma sp.	Aichelberg Dotternhausen	Specimens	A sea sponge (Glass sponge) of the family Staurodermatidae inside Hexactinellida. Found only on depth Basinal deposits, with a funnel like morphology attaining a diameter of at least 15 cm, with the exception of a large specimen of 30 cm diameter found on Dotternhausen.	Stauroderma, a 30 cm diameter specimen from Dotternhausen

Annelida

Genus	Species	Location	Material	Notes	Images
Serpula?[68]	Serpula sp.	Banz Aichelberg Dotternhausen Holzmaden Ohmden Maurach Hemmikon	Multiple Specimens.	A sessile, marine annelid tube worm of the family Serpulidae. Its affinities with the genus Serpula are controversial, since the genus is known mostly since Creataceus strata. Although there are other fossils assigned to the genus on same age deposits of France.[69]	Head of a modern Serpula vermicularis
Filogranula[70][71]	Filogranula tricristata	Banz Aichelberg Dotternhausen Holzmaden Ohmden Maurach Hemmikon	Multiple Specimens.	A sessile, marine annelid tube worm of the family Serpulidae
Tetraserpula[72]	Tetraserpula quadrisulcata	Banz Aichelberg Dotternhausen Holzmaden Ohmden Maurach Hemmikon	Multiple Specimens.	A sessile, marine annelid tube worm of the family Serpulidae.
Mucroserpula[73][74][75]	Mucroserpula quinquecristata	Banz Dotternhausen Holzmaden Ohmden Hemmikon	Multiple Specimens.	A sessile, marine annelid tube worm of the family Serpulidae. It show the characteristic features to live on soft mud ground.
Cementula[73][74][75]	Cementula complanata	Banz Dotternhausen Holzmaden Ohmden Hemmikon	Multiple Specimens.	A sessile, marine annelid tube worm of the family Serpulidae. It show the characteristic features to live on soft mud ground.
Pentaditrupa[73][74][75]	Pentaditrupa quinquesulcata Pentaditrupa cylindracea	Banz Dotternhausen Holzmaden Ohmden Hemmikon	Multiple Specimens.	A sessile, marine annelid tube worm of the family Serpulidae. Pentaditrupa managed to lie freely on the mud, as shows that its tube's curvature provides stability to its position.
Propomatoceros[73][74][75]	Propomatoceros segmentata	Banz Dotternhausen Holzmaden Ohmden Hemmikon	Multiple Specimens.	A sessile, marine annelid tube worm of the family Serpulidae. Denominated "Serpula" segmentata, it lacks the two longitudinal edges characteristic on Mucroserpula.
Glomerula[76]	Glomerula plexus	Banz Altdorf Dotternhausen	Multiple Specimens.	A polychaete worm of the family Sabellidae.	Example of Glomerula specimen
Dictyothylakos[77]	Dictyothylakos pesslerae Dictyothylakos sp.	Irlbach	Leech cocoons	Hirudinea cocoons, identified with palynological residues. The cocoons Dictyothylakos are common on flooded basin sediments, and implies not only the presence of parasitic leeches, but also the presence of large hosts nearby.	Example of Leech Cocoon

Mollusca

Bivalvia

Genus	Species	Location	Material	Notes	Images
Meleagrinella[78][79][80]	Meleagrinella substriata	All the Formation[81]	Thousands of Specimens.	An oxytomid scallop.	Colony of specimens
Oxytoma[78][79]	Oxytoma inequivalvis	All the Formation	Thousands of Specimens.	An oxytomid scallop.
Chlamys[78][79][81][82][80]	Chlamys textoria	All the Formation	Thousands of Specimens.	A pectinoid scallop.	Single specimen
Eopecten[78][79][81][82][80]	Eopecten velatus	All the Formation	Thousands of Specimens.	A pectinoid scallop.
Propeamussium[78][79][81][82][80]	Propeamussium nonarium Propeamusium pumilus	All the Formation	Thousands of Specimens.	A propeamussiid mud scallop.	Various specimens on the same rock
Plicatula[78][79][81][82][80]	Plicatula spinosa	All the Formation	Thousands of Specimens.	A plicatulid mud scallop.
Bositra[82][78][79][81][82][80][83][84]	Bositra buchii	All the Formation	20.000 specimens/m²	A "posidoniid" ostreoidan. It is the type fossil of the Posidonia Shale. Originally it was named "Posidonia bronni", thought to be a new genus, and the strata was denominated the Posidonia layers after it. Years later it turned out to be a junior synonym of Bositra, and thus, it was reassigned. However, the name of the layers was retained. The habitat and mode of life of Bositra has been debated for more than a century. There have been different interpretations, such as a pseudoplanktonic organism,[85] a benthic organism[86] related to open marine floor, where it was the main inhabitant of the basinal settings,[87][88] a free swimming mode of life filtering phytoplankton,[84] and a hybrid mode, where it has a life cycle with holopelagic reproduction controlled by the change on Oxygen levels,[89] and even a chemosymbiotic lifestile, related to the large crinoid rafts, being the main "Safe conduct" to evade anoxic events.[90] All the opinions along the years led to a large study in 1998, where the size/frequency distribution, the density of growth thanks to the lines related to the shell size and the position of the redox boundary by total organic carbon diagrams has revealed that Bositra probably had a benthic mode of life.[91]	Thousands of specimens in one matrix
Steinmannia[78][79][80][85]	Steinmannia bronni Steinmannia radiata	All the Formation	Thousands of Specimens.	A "posidoniid" ostreoidan. Another Genera mistaken with "Posidonia bronni".	Various specimens in one matrix
Gervillella[78][79][81][82][80][83][85]	Gervillella lanceolata	All the Formation	Thousands of Specimens.	A bakevelliid mud oyster.
Liostrea[78][79][81][82][80][83][85]	Liostrea falcifera	All the Formation	Thousands of Specimens.	A gryphaeid mud oyster.	Various specimens
Gryphaea[78][79][81][82][80]	Gryphaea arcuata	All the Formation	Thousands of Specimens.	A gryphaeid mud oyster.	Various specimens
Pseudomytiloides[78][79][81][82][80]	Pseudomonotis substriata Pseudomytiloides dubius	All the Formation	Thousands of Specimens.	An inoceramid clam. Being the second most common genera of Bivalve on the Formation, it had been object to several studies to find its ecological niche, like Bositra. Several opinions include a pseudoplanktonic-only organism, able to live in open sea,[88] or a benthonic-only organism.[87] On the 1998 evaluation with Bositra, was found that probably has a benthic early life that translated to a faculatively pseudoplanktonic mode of adult life.[91]	Single specimen
Nicaniella[78][79][81][82][80]	Nicaniella (Nicaniella) pumila Nicaniella striatosulcata	All the Formation	Thousands of Specimens.	An astartid clam.
Palaeonucula[78][79][81][82][80]	Palaeonucula ungulella	All the Formation	Thousands of Specimens.	A nuculidae nut clam.
Cucullaea[78][79][81][82][80][83]	Cucculea galathea	All the Formation	Thousands of Specimens.	A cucullaeid clam.
Mesomiltha[78][79][81][82][80][83]	Mesomiltha pumila Mesomiltha pulchra	All the Formation	Thousands of Specimens.	A lucinid clam.
Goniomya[78][79][81][82][80][85]	Goniomya rhombifera	All the Formation	Thousands of Specimens.	A pholadomyid clam.
Unicardium[78][79][81][82][80][83]	Unicardium bollense	All the Formation	Thousands of Specimens.	A mactromyid clam.
Pleuromya[78][79][81][82][80]	Pleuromya costata Pleuromya ovata Pleuromya unioides Pleuromya uniformis	All the Formation	Thousands of Specimens.	A pleuromyid clam.	Pleuromya uniformis, two specimens
Plagiostoma[78][79][81][82][80][27]	Plagiostoma giganteum	All the Formation	Thousands of Specimens.	A Limidae clam.	Plagiostoma giganteum, specimen multiview
Gibbirynchia[78][79][81][82][80][83]	Gibbirynchia amalthei	All the Formation	Thousands of Specimens.	A pennospiriferinid rhynchonellatan.

Gastropoda

Genus	Species	Location	Material	Notes	Images
Coelodiscus[92][80]	Coelodiscus minutus	Banz Altdorf Mistelgau Aichelberg Schandelah Dotternhausen Holzmaden Ohmden Hemmikon Dudelange-Zoufftgen	Docens of Specimens.	A Coelodiscidae sea Snail. The is the oldest known holoplanktonic gastropod, thanks to a bilateral symmetrical shells as an adaption to active swimming. Also the most common of the sea snails of the Formation, it is also one of the most varied in size terms, with some of the biggest specimens of snail from the Lower Toarcian know.[92] It has been related to large floating driftwood as one of the primary settlers.[92]
Tatediscus[93]	Tatediscus aratus	Banz Altdorf Mistelgau Aichelberg Schandelah Holzmaden Ohmden	Docens of Specimens.	A Coelodiscidae sea Snail. Possible holoplanktonic gastropod.[92]
Procerithium[78][79][81][82][80]	Procerithium brandi	Banz Altdorf Mistelgau Aichelberg Schandelah Niedersachsen	Docens of Specimens.	A Procerithiidae sea Snail.
Toarctocera[94]	Toarctocera subpunctata	Banz Altdorf Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen	Docens of Specimens.	An Aporrhaidae sea Snail. Among the latest described from the formation, is one of the earliest certain aporrhaidae. Characterised by large spines growing at the head of the cunch.
Cryptaulax[78][79][81][82][80]	Cryptaulax armatum	Banz Altdorf Mistelgau Aichelberg Schandelah Dotternhausen	Docens of Specimens.	A Cryptaulacidae sea Snail.
Neritaria[78][79][81][82][80]	Neritaria papilio	Banz Altdorf Mistelgau Aichelberg Schandelah Niedersachsen Holzmaden Ohmden Hemmikon Dudelange-Zoufftgen	Docens of Specimens.	A Neritariidae sea Snail.	Single specimen
Zygopleura[78][79][81][82][80]	Zygopleura septemcincta	Banz Altdorf Mistelgau Aichelberg Holzmaden Ohmden Hemmikon Dudelange-Zoufftgen	Docens of Specimens.	A Zygopleuridae sea Snail.	Specimens
Levipleura[78][79][81][82][80]	Levipleura blainvillei	Banz Mistelgau Aichelberg Schandelah Niedersachsen Holzmaden Ohmden Dotternhausen Hemmikon	Docens of Specimens.	A Zygopleuridae sea Snail.
Trochus[78][79][81][82][80]	Trochus subduplicatus	Banz Altdorf Mistelgau Aichelberg Schandelah Niedersachsen Holzmaden Ohmden Hemmikon Dudelange-Zoufftgen	Docens of Specimens.	A Trochidae sea Snail.	Specimens
Tylotrochus[78][79][93]	Tylotrochus subimbricatus	Banz Altdorf Mistelgau Aichelberg Holzmaden Ohmden Dotternhausen Hemmikon	Docens of Specimens.	A Trochidae sea Snail. Characterised by a rhomboid scaly pattern on the teleoconch whorls.
Pleurotomaria[78][79][93]	Pleurotomaria debuchii	Banz Altdorf Mistelgau Aichelberg Schandelah Niedersachsen Holzmaden Ohmden Dotternhausen Dudelange-Zoufftgen	Docens of Specimens.	A Pleurotomariidae sea Snail.	Specimen
Ptychomphalus[78][79][93]	Ptychomphalus expansus	Banz Altdorf Mistelgau Aichelberg Schandelah Dotternhausen	Docens of Specimens.	An Eotomariidae sea Snail.
Pterotrachea[95]	Pterotrachea liassica Pterotrachea ceratophagus	Holzmaden Ohmden Aldortf Dotternhausen	Various specimens.	A Pterotracheidae sea Slug. Among the oldest pelagic floating Slugs, Pterotrachea liassica had a more extended larval period than modern extant Pterotrachea coronata, because one additional whorl is Present.[95]	Modern Specimen

Cephalopoda

Genus	Species	Location	Material	Notes	Images
Cenoceras[96][97][98]	Cenoceras intermedium Cenoceras astacoides	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Irlbach Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Nautilidae Nautilidan. Includes the largest specimen of Cenoceras known, with 80 cm width.	Nautilidae shell from Banz, probably Cenoceras
Lytoceras[99][97][98]	Lytoceras ceratophagum Lytoceras onychograptum Lytoceras cornucopia Lytoceras sublineatum Lytoceras germaini Lytoceras mucronatum	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Irlbach Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Lytoceratidae Ammonite. Lytoceras can get quite big, with nearly 50 cm in diameter.
Pachylytoceras[99][97]	Pachylytoceras hircinum Pachylytoceras cf. hircinum Pachylytoceras torulosum Pachylytoceras wrighti Pachylytoceras dilucidum	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Lytoceratidae Ammonite.
Erycites[99][97]	Erycites labrosus	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hammatoceratidae Ammonite.
Hammatoceras[99][97][98]	Hammatoceras insigne Hammatoceras compressum	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Irlbach Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hammatoceratidae Ammonite.	Hammatoceras
Grammoceras[100][98]	Grammoceras thouarsense Grammoceras orhignyi Grammoceras doerntense Grammoceras dispatisum Grammoceras saemanni	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Irlbach Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hildoceratidae Ammonite.	Grammoceras specimen from Banz
Phlyseogrammoceras[100][97]	Phlyseogrammoceras dispansum Phlyseogrammoceras cf. dispansiforme	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hildoceratidae Ammonite.
Pseudogrammoceras[100]	Pseudogrammoceras bingmanni	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hildoceratidae Ammonite.
Hudlestonia[97]	Hudlestonia serrodens Hudlestonia affinis	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hildoceratidae Ammonite.
Catulloceras[97]	Catulloceras dumortieri	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hildoceratidae Ammonite.
Dumortieria[97]	Dumortieria levesquei Dumortieria moorei Dumortieria radians Dumortieria costula Dumortieria cf. costula Dumortieria kochi Dumortieria cf. rustica Dumortieria sparsicosta Dumortieria brancoi Dumortieria tabulata Dumortieria striatulocostata Dumortieria pseudoradiosa Dumortieria radiosa Dumortieria aff. gundershofensis Dumortieria cf. prisca	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hildoceratidae Ammonite.
Cotteswoldia[97]	Cotteswoldia distans Cotteswoldia lotharingica Cotteswoldia mactra Cotteswoldia subcompta Cotteswoldia fluitans	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hildoceratidae Ammonite.
Harpoceras[101][102]	Harpoceras falciferum Harpoceras nitescens	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hildoceratidae Ammonite.	Harpoceras specimen
Tiltoniceras[100]	Tiltoniceras antiquum	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Irlbach Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hildoceratidae Ammonite.	Tiltoniceras specimen
Hildoceras[103][104][98]	Hildoceras levisoni Hildoceras serpentinum[103] Hildoceras semipolitum Hildoceras bifrons	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Irlbach Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hildoceratidae Ammonite.	Hildoceras specimen
Mercaticeras[103]	Mercaticeras forte	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hildoceratidae Ammonite.	Mercaticeras specimen
Eleganticeras[103]	Eleganticeras exaratum	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hildoceratidae Ammonite.
Pseudolioceras[103][104]	Pseudolioceras compactile Pseudolioceras discoides	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Hildoceratidae Ammonite.
Dactylioceras[103][105][98]	Dactylioceras semiannulatum Dactylioceras vermis Dactylioceras athleticum Dactylioceras annulatum	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Irlbach Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Dactylioceratidae Ammonite.	Dactylioceras commune on Holzmaden
Catacoeloceras[104][97][98]	Catacoeloceras crassum Catacoeloceras raquinianum	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Dactylioceratidae Ammonite.
Coeloceras[104][97][98]	Coeloceras crassum Coeloceras mucronatum	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	Type Coeloceratidae Ammonite.
Phylloceras[103][106]	Phylloceras heterophyllum Phylloceras supraliasicum	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Phylloceratidae Ammonite. The largest ammonite found in the Posidonienschiefer comes from the Ohmden quarry,and belongs to a Phylloceras heterophyllum with a diameter of 87 cm.[103]	Phylloceras restoration
Haugia[103][104]	Haugia variabilis Haugia illustris Haugia jugosa	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Phymatoceratidae Ammonite.
Denckmannia[103][104]	Denckmannia malagma	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Phymatoceratidae Ammonite.
Phymatoceras[103][104]	Phymatoceras lilli	Dudelange-Zoufftgen	Multiple Specimens.	A Phymatoceratidae Ammonite.
Loligosepia[107]	Loligosepia bucklandi	Banz Dotternhausen Holzmaden Ohmden Aselfingen	Multiple Specimens.	A Loligosepiidae Loligosepiidan.[108] The Loligosepiidae is believed to be ancestral to the Recent vampire squid, Vampyroteuthis infernalis.[109]	Loligosepia Holzmaden specimen
Jeletzkyteuthis[110]	Jeletzkyteuthis coriaceus	Holzmaden Ohmden Banz Dotternhausen	Multiple Specimens.	A Loligosepiidae Loligosepiidan. Related to the modern Vampyroteuthis infernalis.
Geopeltis[111]	Geopeltis simplex	Holzmaden Ohmden Banz	Multiple Specimens.	A Loligosepiidae Loligosepiidan. Related to the modern Vampyroteuthis infernalis.	Geopeltis specimen
Parabelopeltis[112]	Parabelopeltis flexuosa	Holzmaden Ohmden Banz Altdorf	Multiple Specimens.	A Loligosepiidae Loligosepiidan. Related to the modern Vampyroteuthis infernalis.
Chitinobelus[113]	Chitinobelus acifer	Holzmaden Ohmden	Multiple Specimens.	A Belemnotheutidae Belemnite. Chitinobelus is an extrange fossil, sice the rostrum was composed of aragonite with organic material, while normal Belemnites had calcite. Has been suggested this rostrum was calcitic.
Chondroteuthis[114]	Chondroteuthis wunnenbergi	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Belemnotheutidae Belemnite.
Simpsonibelus[104]	Simpsonibelus dorsalis Simpsonibelus lentus[115]	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Belemnotheutidae Belemnite.
Acrocoelites[103][104]	Acrocoelites ilminstrensis Acrocoelites pyramidalis Acrocoelites glaber Acrocoelites cf. riegrafi Acrocoelites levidensis Acrocoelites vulgaris Acrocoelites tripartitus	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Megateuthididae Belemnite. Includes some of the Biggest Know Belemnites, with stimated maximum up to 4.5 m long in life, altrougth, most specimens would have been rather smaller.
Dactyloteuthis[103][104]	Dactyloteuthis wrighti Dactyloteuthis digitalis[115] Dactyloteuthis semistriata Dactyloteuthis irregularis Dactyloteuthis similis Dactyloteuthis incurvata	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Megateuthididae Belemnite.
Youngibelus[116][117]	Youngibelus giganteus Youngibelus ohmdenensis[115] Youngibelus simpsoni	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Megateuthididae Belemnite. Includes really large specimens	Youngibelus Reconstruction
Passaloteuthis[118]	Passaloteuthis paxillosa Passaloteuthis bisulcata	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Passaloteuthididae Belemnite.	Passaloteuthis Holzmaden specimen
Salpingoteuthis[115]	Salpingoteuthis trisulcata Salpingoteuthis bauhini Salpingoteuthis longisulcata Salpingoteuthis macra Salpingoteuthis tessoniana Salpingoteuthis dorsetiensis Salpingoteuthis blomenhofensis	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple Specimens.	A Salpingoteuthididae Belemnite.	Salpingoteuthis specimen
Clarkeiteuthis[119]	Clarkeiteuthis conocauda	Banz Aichelberg Dotternhausen Holzmaden Ohmden	Multiple Specimens.	A Diplobelidae Coleoidean. It has been found adult individuals of Clarkeiteuthis which caught small teleost fish of the species Leptolepis bronni. Further indirect evidence for the hunting behaviour comes from their body orientation in the water during life.[120]	Clarkeiteuthis Holzmaden specimen
Lioteuthis[121]	Lioteuthis problematica	Holzmaden Ohmden Dotternhausen	Various Specimens.	Type member of the Lioteuthididae Squid family. The taxonomic position of Lioteuthis is uncertain, although the wings reaching the proximal gladius section and the smooth median field suggest affinity to the Prototeuthididae[121]
Teudopsis[122]	Teudopsis bollensis Teudopsis subcostata Teudopsis schubleri	Banz Aichelberg Dotternhausen Holzmaden Ohmden Aselfingen	Various Specimens.	A Teudopseina Squid.	Teudopsis Ohmden specimen
Geotheutis[123]	Geotheutis bollensis	Dotternhausen Holzmaden Ohmden	Various Specimens.	A possible primigenial Cuttlefish. Is one of the most important fossils of Cephalopods on the Posidonia Shale, due to be one of the Earliest examples of Pigments found on any species, also one of the first historically.[124] The pigments are preserved on various specimens with Eumelanin related to its ink sacs and include even phosphatized musculature.[125]

Arthropoda

Cycloidea

Genus	Species	Location	Material	Notes	Images
Juracyclus[126]	Juracyclus posidoniae	Tübingen	Partial Specimens.	The First Cycloid Arthropod from the Jurassic, from the family Cycloidae inside Cycloidea.[126] Cycloids are a group of maxillopod arthropods that span between the Paleozoic until the latest Cretaceous, probably related to the crustaceans and probably detritivores.[126]

Malacostraca

Genus	Species	Location	Material	Notes	Images
Orhomalus[127]	Orhomalus arpi	Neumarkt	Partial Specimens.	A hermit crab of the family Paguroidea. Hermit crabs are common of the marine layers of the lower Jurassic Europe, mostly on the Pliensbachian-Toarcian Boundary of France and Germany. Orthomalus is a small sized crab, probably inhabitant of nearshore waters, and even deltaic and beach environments, similar to modern genera.
Palaeopagurus[128]	Palaeopagurus sp.	Holzmaden	Single Specimen inside an Ammonite Shell.	An hermit crab of the family Paguridae.
Uncina[129][130]	Uncina posidoniae	Holzmaden Ohmden Dotternhausen Banz Mistlegau Altdorf	Nearly complete & Partial Specimens.	An Astacidea Decapodan of the family Uncinidae. Reaching large sizes of almost half a meter (39–47 cm), Uncina Posidoniae is among the largest know Jurassic Crustaceans. Uncina posidoniae is also the largest representative of the genus Uncina.[130] This large crustacean has been found associated with Ammonite and Bivalve filled Bentos, where probably hunted different kinds of prey.[130] Its large claws would have been perfect for hunt small invertebrates and vertebrates.[130]	Uncina posidoniae specimen The Largest complete Uncina posidoniae specimen, with 44 cm long.
Tonneleryon[131]	Tonneleryon schweigerti	Holzmaden Ohmden Dotternhausen	Partial Specimens and complete Specimens	A gregarious Polychelidan Lobster.
Proeryon[132][133]	Proeryon giganteus Proeryon hauffi[134] Proeryon hartmanni[135] Proeryon laticaudatus[136]	Holzmaden Ohmden Dotternhausen Banz Mistlegau Altdorf	Partial Specimens.	A Coleiidae Decapodan. The largest Coeloid from the formation, P. giganteus is a species reaching a larger size than most other polychelidans, with up to 15 cm. On the Posidonia Shale there is the most abundant variety of species from the genus, ranging from different sizes and morphologies, that indicate different habitat & feeding adaptations on the genus. Some like P. hartmani show less adaptations to hunt for small nectobenthic preys than other relatives, being abundant on Oyster-filled waters. There is a relatively abundance of the genus in deep-water settings from the Toarcian onward.[137]	Proeryon hartmanni specimen from Banz Proeryon sp. specimen from Schandelah
Antrimpos[138][139]	Antrimpos noricus	Holzmaden Ohmden Dotternhausen Banz	Partial Specimens.	A Penaeidae Decapodan.	Antrimpos specimen
Palaeastacus[140][141]	Palaeastacus sp.	Holzmaden Ohmden Dotternhausen	Partial Specimens.	A Erymidae Decapodan.	Palaeastacus specimen
Coleia[142]	Coleia theodorii Coleia moorei Coleia sinuata	Holzmaden Ohmden Dotternhausen Banz	Complete Specimens. Includes specimens inside ammonites shells	An Erymidae Decapodan.
Eryma[143][144][145]	Eryma amalthei Eryma sp.	Holzmaden Ohmden Dotternhausen Banz	Complete Specimens	Type genus of the Erymidae Decapodan family. Originally, was named Glyphea amalthei,informally used by Quenstedt and housed on the Museum Naturkunde in Württemberg. A series of posterior revisions probe it was a different genus.[146]	Eryma specimen
Stenodactylina[147][148]	Stenodactylina liasina	Holzmaden Ohmden Dotternhausen	Complete Specimens	An Erymidae Decapodan.
Achelata[149]	Achelata gen. et sp. indet.	Holzmaden	Single complete specimen in late larval stage	The specimen reported represents the oldest fossil record of an achelatan lobster larva, and the first representative of achelatan lobsters in the Posidinia Shale. Shares similarities with the late jurassic genus Cancrinos.	California spiny lobsters are relatives of the Larval specimen from the Posidonia Shale
Stomatopoda[150]	Stomatopoda Indeterminate	Holzmaden	Single Incomplete Specimen	A Stomatopoda Malacostracan. Its affinities haven't been tested.	A female Odontodactylus Scyllarus mantis shrimp, maybe related to the Posidonia Specimen

Thoracica

Genus	Species	Location	Material	Notes	Images
Toarcolepas[151]	Toarcolepas mutans	Aichelberg	Numerous disarticulated individuals, associated with fossil wood.[151]	A phospatic-shelled Cirripede of the family Eolepadidae.[151] Toarcolepas is provisionally interpreted as the oldest epiplanktonic cirripede known, and is thought to have lived attached to floating driftwood.[151]	Modern genus Lepas is the most related taxa to Toarcolepas

Arachnida

Genus	Species	Location	Material	Notes	Images
Liassoscorpionides[152]	Liassoscorpionides schmidti	Niedersachsen	Single Incomplete Specimen.	Being the only Jurassic scorpion known, there is no evidence that L. schmidti was aquatic (which was suggested in the past) and in the absence of further, better preserved material it should be excluded from future considerations of broad patterns of scorpion evolution.[153]	Speculative Reconstruction

Insecta

Insects are a common terrestrial animals that where proabaly drifted to the sea due to Moonsonal conditions present on the Posidonia Shale.[154]

Genus	Species	Location	Material	Notes
Heterophlebia[155][156]	Heterophlebia buckmani	Holzmaden Schandelah Braunschweig Hattorf, Fallersleben Bascharage	Specimens	A Dragonfly of the family Heterophlebiidae. A relative abundant genus, present on most of the pits of the Posidonia Shale, even on Holzmaden.
Plagiophlebia[156]	Plagiophlebia praecostarea	Hattorf, Fallersleben	Specimens	A Dragonfly of the family Heterophlebiidae.
Heterothemis[157]	Heterothemis brodiei	Holzmaden Schandelah Beienrode Hattorf, Fallersleben Hemmikon[158] Bascharage	Specimens	A Dragonfly of the family Liassogomphidae. Second most abundant genus of Odonatan in the Formation.
Phthitogomphus[156][159]	Phthitogomphus angulatus	Niedersachsen Luxguard quarry	Specimens	A Dragonfly of the family Liassogomphidae.
Proinogomphus[156]	Proinogomphus bodei Proinogomphus kreuzerorum[160]	Hattorf, Fallersleben	Specimens	A Dragonfly of the family Liassogomphidae.
Ensphingophlebia[156]	Ensphingophlebia undulata	Braunschweig	Specimens	A Dragonfly of the family Sphenophlebiidae.
Mesoepiophlebia[159]	Mesoepiophlebia veronicae	Bascharage	Specimens	A Dragonfly of the family Sphenophlebiidae.
Liassostenophlebia[156]	Liassostenophlebia germanica	Rhine-Danube canal, Km 112	Specimens	A Dragonfly
Syrrhoe[156]	Syrrhoe commissa	Niedersachsen	Specimens	A Dragonfly.
Strongylogomphus[156]	Strongylogomphus grasselianus	Niedersachsen	Specimens	A Dragonfly.
Myopophlebia[156][159]	Myopophlebia libera	Beienrode Bascharage	Specimens	A Dragonfly of the family Myopophlebiidae.
Paraheterophlebia[156][159]	Paraheterophlebia wunnenbergi Paraheterophlebia marcusi	Niedersachsen Bascharage	Specimens	A Dragonfly of the family Myopophlebiidae.
Paraplagiophlebia[159]	Paraplagiophlebia loneuxi	Bascharage	Specimens	A Dragonfly of the family Myopophlebiidae.
Necrogomphus[156][161]	Necrogomphus brunswigae	Schandelah	Specimens	A Dragonfly of the Family Liassogomphidae.
Campterophlebia[156][161]	Campterophlebia elegans	Schandelah	Specimens	A Dragonfly of the family Campterophlebiidae.The largest Early Jurassic Insect Know, with a wings size up to 20 cm.[162]
Gallodorsettia[163]	Gallodorsettia kronzi	South of Foetz	Specimens	A Dragonfly of the family Campterophlebiidae.
Henrotayia[164]	Henrotayia marci	Bascharage	Specimens	A Dragonfly of the family Henrotayiidae.
Liadoblattina[165]	Liadoblattina blakei	Holzmaden Niedersachsen	Specimens	A Cockroach of the family Raphidiomimidae.
Ptyctoblattina[156]	Ptyctoblattina simplicior Ptyctoblattina dilatata	Niedersachsen Beienrode	Specimens	A Cockroach of the family Raphidiomimidae.
Caloblattina[156]	Caloblattina mathildae	Niedersachsen Beienrode	Specimens	A Cockroach of the family Caloblattinidae.
Blattula[156]	Blattula langfeldti	Niedersachsen Beienrode	Specimens	A Cockroach of the family Blattulidae.
Schesslitziella[166][167][168]	Schesslitziella haupti Schesslitziella integra	Feuermühlenberg near Scheßlitz Kerkhofen	Specimens	A Stick insect. One of the described insects found more near the Bohemian Massif, where probably belong most of the terrestrial invertebrate fauna.
Chresmodella	Chresmodella fissa	Hattorf, Fallersleben	Specimens	A Stick insect of the family Aerophasmidae.
Compactofulgoridium[156]	Compactofulgoridium spoliatum	Niedersachsen	Specimens	A Grasshopper.
Protogryllus[156]	Protogryllus formosus Protogryllus hattorfensis Protogryllus praeacutus Protogryllus symmetricus Protogryllus multoramosus Protogryllus multovenosus Protogryllus laceratus Protogryllus foliolum Protogryllus implicatus Protogryllus fissus	Niedersachsen	Specimens	A Grasshopper of the family Protogryllidae.
Panorpidium[156]	Panorpidium media Panorpidium geinitzi Panorpidium minima	Niedersachsen	Specimens	A Grasshopper of the family Elcanidae.
Acridiopsis[156]	Acridiopsis spoliata	Hattorf, Fallersleben	Specimens	A Short-horned Grasshopper of the family Acrididae.
Locustopsis[156]	Locustopsis procera Locustopsis bernstorffi Locustopsis maculosa	Niedersachsen	Specimens	A Grasshopper of the family Locustopsidae.
Liadolocusta[156]	Liadolocusta ornata	Niedersachsen	Specimens	A Grasshopper of the family Locustopsidae.
Archijassus[156]	Archijassus heeri	Niedersachsen	Specimens	A Planthopper of the family Archijassidae.
Elasmoscelidium[156]	Elasmoscelidium rectemarginatum Elasmoscelidium promotum Elasmoscelidium boreale	Niedersachsen Braunschweig	Specimens	A Planthopper.
Fulgoridium[156]	Fulgoridium mancomarginatum Fulgoridium semiperspicuum Fulgoridium cubitoramosum Fulgoridium cuneiforme Fulgoridium infuscatum Fulgoridium paulodilatatum Fulgoridium exiguemaculatum Fulgoridium reduncum Fulgoridium fallerslebense Fulgoridium hattorfense Fulgoridium gottingense Fulgoridium tenuimaculatum Fulgoridium incurvatum Fulgoridium praeobtusum Fulgoridium raromaculatum Fulgoridium cubitofurcatum Fulgoridium basilaesum Fulgoridium hondelanum Fulgoridium fabri Fulgoridium hildesheimense Fulgoridium symmetricum Fulgoridium latius Fulgoridium posidonicum Fulgoridium silvaticum	Niedersachsen Flechtorf near Fallersleben Aselfingen	Specimens	A Planthopper of the family Fulgoridiidae.
Fulgoridulum[156]	Fulgoridulum egens	Niedersachsen	Specimens	A Planthopper of the family Fulgoridiidae.
Procerofulgoridium[156]	Procerofulgoridium verticillatum Procerofulgoridium praefastigatum	Niedersachsen	Specimens	A Planthopper of the family Fulgoridiidae.
Tetrafulgoria[156]	Tetrafulgoria parallelogramma	Niedersachsen	Specimens	A Planthopper of the family Fulgoridiidae.
Metafulgoridium[156]	Metafulgoridium praetruncatum Metafulgoridium spatulaeforme	Niedersachsen	Specimens	A Planthopper of the family Fulgoridiidae.
Productofulgoridium[156]	Productofulgoridium filiferum Productofulgoridium praeacutum	Niedersachsen	Specimens	A Planthopper of the family Fulgoridiidae.
Margaroptilon[156]	Margaroptilon formosum Margaroptilon cuneatum Margaroptilon paucisinuatum Margaroptilon detruncatum Margaroptilon procerum	Niedersachsen	Specimens	A Planthopper of the family Fulgoridiidae.
Compactofulgoridium[156]	Compactofulgoridium fronterotundum Compactofulgoridium obesum Compactofulgoridium aries Compactofulgoridium decapitatum Compactofulgoridium concameratum Compactofulgoridium paenintegrum	Niedersachsen	Specimens	A Planthopper of the family Fulgoridiidae.
Procercopis[156]	Procercopis lacerata Procercopis abscissa Procercopis wunnenbergi	Niedersachsen	Specimens	A Froghopper of the family Procercopidae.
Megalocoris[156]	Megalocoris laticlavus	Niedersachsen	Specimens	A Shore bug.
Eurynotis[156]	Eurynotis incisus	Niedersachsen	Specimens	A Shore bug of the family Archegocimicidae.
Somatocoris[156]	Somatocoris conservatus	Niedersachsen	Specimens	A Shore bug of the family Archegocimicidae.
Corynecoris[156]	Corynecoris occultatus Corynecoris semigranulatus	Niedersachsen	Specimens	A Shore bug of the family Archegocimicidae.
Entomecoris[156]	Entomecoris minor	Niedersachsen	Specimens	A Shore bug of the family Archegocimicidae.
Ensphingocoris[156]	Ensphingocoris parvulus Ensphingocoris praerotundatus	Niedersachsen	Specimens	A Shore bug of the family Archegocimicidae.
Engynabis[156]	Engynabis tenuis	Niedersachsen	Specimens	A Shore bug of the family Archegocimicidae.
Apicasia[156]	Apicasia inolata	Niedersachsen	Specimens	A basal Beetle.
Pholipheron[156]	Pholipheron articulatus	Niedersachsen	Specimens	A basal Beetle.
Grasselites[156]	Grasselites pusillus	Niedersachsen	Specimens	A basal Beetle.
Omogongylus[156]	Omogongylus ovatus	Niedersachsen	Specimens	A basal Beetle.
Sideriosemion[156]	Sideriosemion punctolineatum	Niedersachsen	Specimens	A basal Beetle.
Metanastes[156]	Metanastes denudatus	Niedersachsen	Specimens	A basal Beetle.
Diatrypamene[156]	Diatrypamene angulocollis	Niedersachsen	Specimens	A basal Beetle.
Aptilotitus[156]	Aptilotitus capitecarens	Niedersachsen	Specimens	A basal Beetle.
Rhomaleus[156]	Rhomaleus ornatus	Niedersachsen	Specimens	A basal Beetle.
Leptosolenophorus[156]	Leptosolenophorus brevicollis	Niedersachsen	Specimens	A basal Beetle.
Gastroratus[156]	Gastroratus dispertitus	Niedersachsen	Specimens	A basal Beetle.
Brachylaimon[156]	Brachylaimon striatus	Niedersachsen	Specimens	A basal Beetle.
Pleuralocista[156]	Pleuralocista insculpta	Niedersachsen	Specimens	A basal Beetle.
Mesoncus[156]	Mesoncus striatulus	Niedersachsen	Specimens	A basal Beetle.
Palaeotrachys[156]	Palaeotrachys laticollis	Niedersachsen	Specimens	A basal Beetle.
Hydroicetes[156]	Hydroicetes affictus	Schandelah	Specimens	A basal Beetle.
Scalopoides[156]	Scalopoides inscissus	Niedersachsen	Specimens	A basal Beetle.
Peridosoma[156]	Peridosoma praecisum	Niedersachsen	Specimens	A basal Beetle.
Syntomopterus[156]	Syntomopterus latus	Beienrode	Specimens	A basal Beetle.
Sphaerocantharis[156]	Sphaerocantharis striata Sphaerocantharis defossa	Beienrode	Specimens	A basal Beetle.
Rhysopsalis[156]	Rhysopsalis distorta	Beienrode	Specimens	A basal Beetle.
Diplocelides[156]	Diplocelides minutus	Niedersachsen	Specimens	A basal Beetle.
Tripsalis[156]	Tripsalis praecisa	Niedersachsen	Specimens	A basal Beetle.
Trochiscites[156]	Trochiscites capitapertus	Niedersachsen	Specimens	A basal Beetle.
Prosynactus[156]	Prosynactus gracilis Prosynactus scissus Prosynactus procerus	Beienrode Niedersachsen	Specimens	A False Ground Beetle of the family Trachypachidae.
Coreoeicos[156]	Coreoeicos dilatatus	Beienrode	Specimens	A False Ground Beetle of the family Trachypachidae.
Aposphinctus[156]	Aposphinctus conservatus Aposphinctus striatus	Beienrode	Specimens	A Water Scavenger Beetle of the family Hydrophilidae.
Zetemenos[156]	Zetemenos sexlineatus	Niedersachsen	Specimens	A basal Beetle.
Amphoxyne[156]	Amphoxyne lineata Amphoxyne minuta	Beienrode	Specimens	A basal Beetle.
Tolype[156]	Tolype rotundata	Beienrode	Specimens	A basal Beetle.
Amblycephalonius[156]	Amblycephalonius tenuistriatus	Niedersachsen	Specimens	A basal Beetle of the family Coptoclavidae.
Ooperioristus[156]	Ooperioristus applanatus	Niedersachsen	Specimens	A basal Beetle of the family Coptoclavidae.
Camaricopterus[156]	Camaricopterus ovalis	Niedersachsen	Specimens	A basal Beetle of the family Schizophoridae.
Megachorites[156]	Megachorites brevicollis	Niedersachsen Volkmarsdorf, Braunschweig	Specimens	A Giant Beetle. It is among the largest found on all the Jurassic.
Protobittacus[156]	Protobittacus desacuminatus	Niedersachsen Hattorf, Fallersleben	Specimens	A Hangingfly of the family Bittacidae.
Parabittacus[156]	Parabittacus lingula	Niedersachsen Hattorf, Fallersleben	Specimens	A Hangingfly of the family Bittacidae.
Haplobittacus[156]	Haplobittacus parvus	Niedersachsen	Specimens	A Hangingfly of the family Bittacidae.
Mesobittacus[156]	Mesobittacus clavaeformis Mesobittacus marginelaesus	Beienrode Niedersachsen	Specimens	A Hangingfly of the family Bittacidae.
Orthophlebia[156]	Orthophlebia latipennisimilis Orthophlebia fallerslebensis Orthophlebia elongata	Niedersachsen Hattorf, Fallersleben	Specimens	A Scorpionfly of the family Orthophlebiidae.
Parorthophlebia[156]	Parorthophlebia grasselensis	Niedersachsen Hattorf, Fallersleben	Specimens	A Scorpionfly of the family Orthophlebiidae.
Mesopanorpa[156]	Mesopanorpa formosa	Beienrode	Specimens	A Scorpionfly of the family Orthophlebiidae.
Pseudopolycentropus[156]	Pseudopolycentropus obtusus	Niedersachsen Hattorf, Fallersleben	Specimens	A Scorpionfly of the family Pseudopolycentropodidae.
Homoeoptychopteris[156]	Homoeoptychopteris incerta	Niedersachsen	Specimens	A Fly.
Amianta[156]	Amianta eurycephala	Niedersachsen	Specimens	A Fly.
Culiciscolex[156]	Culiciscolex gibberatus	Niedersachsen	Specimens	A Fly.
Liassonympha[156]	Liassonympha compacta	Niedersachsen	Specimens	A Fly.
Bodephora[156]	Bodephora arucaeformis	Niedersachsen	Specimens	A Fly.
Apistogrypotes[156]	Apistogrypotes inflexa	Niedersachsen	Specimens	A Fly.
Amphipromeca[156]	Amphipromeca acuta	Niedersachsen	Specimens	A Fly.
Cyrtomides[156]	Cyrtomides maculatus	Niedersachsen	Specimens	A Fly.
Sphallonymphites[156]	Sphallonymphites decurtatus	Niedersachsen	Specimens	A Fly.
Propexis[156]	Propexis incerta	Niedersachsen	Specimens	A Fly.
Archipleciomima[169]	Archipleciomima germanica	Niedersachsen	Specimens	A Fly.
Protoplecia[156]	Protoplecia hattorfensis	Niedersachsen	Specimens	A Fly of the family Protopleciidae.
Mesorhyphus[169]	Mesorhyphus ulrichi	Niedersachsen	Specimens	A Wood Gnat of the family Anisopodidae.
Metaraphidia[156]	Metaraphidia vahldieki	Niedersachsen	Specimens	A Snakefly.
Heterorhyphus[156]	Heterorhyphus analivarius Heterorhyphus anomalus	Niedersachsen	Specimens	A Fly.
Amblylexis[156]	Amblylexis gibberata	Niedersachsen	Specimens	A Fly.
Ellipibodus[156]	Ellipibodus laesa	Niedersachsen	Specimens	A Fly.
Homoeoptychopteris[156]	Homoeoptychopteris incerta	Niedersachsen	Specimens	A Fly.
Protorhyphus[156]	Protorhyphus ovisimilis	Niedersachsen	Specimens	A Fly of the family Protorhyphidae.
Praemacrochile[156]	Praemacrochile decipiens	Niedersachsen	Specimens	A primitive Crane fly of the family Tanyderidae.
Nannotanyderus[167][170]	Nannotanyderus krzeminskii	Kerkhofen	Specimens	A primitive Crane fly of the family Tanyderidae.
Architipula[156]	Architipula bodeisimilis Architipula formosa Architipula basiminuta Architipula robusta Architipula clara Architipula bodei Architipula brunsvicensis Architipula analiramosa Architipula aequabilis Architipula fragmentosa Architipula veris Architipula latealata	Niedersachsen	Specimens	A Crane fly of the family Limoniidae.
Ozotipula[156]	Ozotipula tarda	Niedersachsen	Specimens	A Crane fly of the family Limoniidae.
Haplotipula[156]	Haplotipula majalis Haplotipula cubitoramosa	Niedersachsen	Specimens	A Crane fly of the family Limoniidae.
Leptotipuloides[156]	Leptotipuloides fastigata	Niedersachsen	Specimens	A Crane fly.
Mikrotipula[156]	Mikrotipula dixaeformis	Niedersachsen	Specimens	A Crane fly.
Eoptychoptera[167][171]	Eoptychoptera simplex	Niedersachsen Kerkhofen	Specimens	A Phantom Crane fly of the family Ptychopteridae.
Necrotaulius[167][156][154]	Necrotaulius parvulus	Holzmaden Niedersachsen Kerkhofen Hattorf, Fallersleben	Specimens	A Caddisfly.
Prohemerobius[156][172]	Prohemerobius prodromus Prohemerobius septemvirgatus Prohemerobius mediolatus	Niedersachsen	Specimens	A Lacewing of the family Prohemerobiidae.
Parhemerobius[156]	Parhemerobius dilatatus Parhemerobius bodei	Niedersachsen	Specimens	A Lacewing of the family Prohemerobiidae.
Paractinophlebia[156]	Paractinophlebia grasselensis	Niedersachsen	Specimens	A Lacewing of the family Prohemerobiidae.
Liassopsychops[156]	Liassopsychops curvata	Niedersachsen	Specimens	A Lacewing of the family Prohemerobiidae.
Tetanoptilon[156]	Tetanoptilon brunsvicense	Niedersachsen	Specimens	A lance Lacewing of the family Osmylidae.
Protoaristenymphes[173]	Protoaristenymphes bascharagensis	Bascharage	Specimens	A lance Lacewing of the family Mesochrysopidae.
Liassocicada[156][174][167]	Liassocicada mueckei Liassocicada antecedens	Rhine-Danube canal, Km 112 Beienrode	Specimens	A Hairy Cicada of the family Tettigarctidae.
Liassotettigarcta[167]	Liassotettigarcta mueckei	Kerkhofen	Specimens	A Hairy Cicada of the family Tettigarctidae.
Xulsigia[175]	Xulsigia karetsa	Bommelscheier industrial area	Specimens	A Sternorrhynchan of the family Pincombeomorpha. It is curious for its peculiar venation on its wings. Has been proposed its own family, Xulsigiidae.
Indutionomarus[176]	Indutionomarus treveriorum	Bommelscheier industrial area	Specimens	A Coleorrhynchan of the family Progonocimicidae.
Thilopterus[156]	Thilopterus lampei	Schandelah	Specimens	A Wasp of the family Ephialtitidae.
Symphytopterus[156]	Symphytopterus liasinus	Niedersachsen	Specimens	A Wasp of the family Ephialtitidae.
Liadobracona[156]	Liadobracona raduhna	Niedersachsen	Specimens	A Wasp of the family Ephialtitidae.
Pseudoxyelocerus[177]	Pseudoxyelocerus bascharagensis	Bascharage	Specimens	A Wood Wasp of the family Xyelotomidae.
Agmatozoon[156]	Agmatozoon articulatum	Niedersachsen	Specimens	A non classified Insect.
Campeulites[156]	Campeulites cylindricus	Niedersachsen	Specimens	A non classified Insect.
Tomeferusa[156]	Tomeferusa abdita	Niedersachsen	Specimens	A non classified Insect.
Trimerocephalium[156]	Trimerocephalium incisum	Niedersachsen	Specimens	A non classified Insect.
Elasmoscolex[156]	Elasmoscolex hamatus	Niedersachsen	Specimens	A non classified Insect.
Epimetrophora[156]	Epimetrophora recta	Niedersachsen	Specimens	A non classified Insect.
Oocephalina[156]	Oocephalina mutilata	Niedersachsen	Specimens	A non classified Insect.
Platycorion[156]	Platycorion utroquelaesum	Niedersachsen	Specimens	A non classified Insect.
Protomyrmeleon[156][178]	Protomyrmeleon grasselensis Protomyrmeleon brunonis[179]	Niedersachsen Bascharage	Specimens	A Dicondylian, from the family Protomyrmeleontidae.
Dorniella[156]	Dorniella pulchra	Beienrode	Specimens	A basal Insect of the family Blattogryllidae.
Geinitzia[156][180]	Geinitzia latrunculorum Geinitzia fasciata Geinitzia dorni	Niedersachsen	Specimens	A basal Insect of the family Geinitziidae.
Adelocoris[156]	Adelocoris ambiguus	Niedersachsen	Specimens	A basal Insect of the family Pachymeridiidae.
Stiphroschema[156]	Stiphroschema longealatum	Niedersachsen	Specimens	A basal Insect of the family Pachymeridiidae.

Echinodermata

Asterozoa

Genus	Species	Location	Material	Notes	Images
Ophiopholis[181]	Ophiopholis trispinosa	Holzmaden Ohmden Altforf Banz Mistelau Hemmikon	Few Specimens.	An Ophiuridan of the family Ophiactidae. Very rare on the layers.	Modern Specimen
Mesophiomusium[181]	Mesophiomusium geisingense[182] Mesophiomusium scabrum	Holzmaden Ohmden Altforf Banz Mistelau Hemmikon	Few Specimens.	An Ophiuridan of the family Ophiolepididae. Very rare on the layers, being Mesophiomusium geisingense the most common of the two species present.
Ophiarachna[181]	Ophiarachna liasica	Holzmaden Ohmden Altforf Banz Mistelau Hemmikon	Various Specimens.	An Ophiuridan of the family Ophiacanthida. Very Common, related to non anoxic water sedimentation.
Sinosura[181][183]	Sinosura brodiei	Holzmaden Ohmden	Various Specimens.	An Ophiuridan of the family Aplocomidae. Very Common.	Fossil Specimen
?Ophiura[181]	Ophiura astonensis	Holzmaden Ohmden	Few Specimens.	An Ophiuridan of the family Ophiuridae. Its relationships haven't been confirmed and it is based on very fragmentary remains.	Appearance
?Ophiocten[181]	Ophiocten seeweni	Holzmaden Ohmden	Few Specimens.	An Ophiuridan of the family Ophiuridae. Its relationships haven't been confirmed and it is based on very fragmentary remains.
?Pentasteria[181]	Pentasteria sp.	Banz	Few Specimens.	An Asteroidean of the family Astropectinidae. It is very rare on the layers, and a few fragmentary specimens are known. Only a relatively complete specimen is known from Banz Abbey.	Pentasteria sp. from Banz

Echinoidea

Genus	Species	Location	Material	Notes	Images
Cidaris[184]	Cidaris sp.	Holzmaden Ohmden Altforf Banz Irlbach Hemmikon	Few Specimens.	A sea urchin of the family Cidaridae. Common on several layers. Cidaris is genus that still alive today. A bottom dweller, is commonly found associated with Belemnnite fossils, probably due to eating its carcasses.	Modern Specimen
Diademopsis[99]	Diademopsis crinifera Diademopsis sp.	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Holzmaden Ohmden Irlbach Aselfingen Maurach Hemmikon Bascharage	Docens of Specimens	A sea urchin of the family Pedinidae. It is the most common sea urchin in the formation, present on all the levels with specimens of various sizes. Vinculated to sea bottom sediments, before Gasteropods and Bivalves, Diademospsis was the tird major colonizer of the bottom, in between anoxic changes.	Specimens from Holzmaden
Procidaris[99]	Procidaris sp.	Holzmaden Banz Irlbach	Few Specimens.	A sea urchin of the family Miocidaridae

Holothuroidea

Genus	Species	Location	Material	Notes	Images
Uncinulina[185]	Uncinulina parvispinosa	Maurach	Few Specimens.	A Sea Cucumber of the Family Elasipodida. Common on the Austrian realm, characterised due to been present of Pelagic environments, as is common on modern genera of its family, but not on the fossil record. Can be the oldest pelagic representative of the Sea Cucumber family. Can be also related to the modern deep sea genus Enypniastes.	Modern Species Enypniastes can be related to Uncinulina

Crinoidea

Genus	Species	Location	Material	Notes	Images
Praetetracrinus[186]	Praetetracrinus kutscheri	Banz Altdorf Holzmaden Ohmden	Multiple specimens.	A Crinoidean of the family Plicatocrinidae.
Shroshaecrinus[186]	?Shroshaecrinus quedlinburgensis	Banz Altdorf Holzmaden Ohmden	Multiple specimens.	A Crinoidean of the family Millericrinidae.
Procomaster[181]	Procomaster pentadactylus	Banz Altdorf Holzmaden Ohmden	Multiple specimens.	A Crinoidean of the family Isocrinida.
Isocrinus[187]	Isocrinus basaltiformis	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Dotternhausen Holzmaden Ohmden Maurach	Multiple specimens.	Type genus of Crinoidean of the family Isocrinida.	Close view of one specimen Single specimen
Seirocrinus[40]	Seirocrinus subangularis	Banz Altdorf Mistelgau Aichelberg Schandelah Dotternhausen Holzmaden Ohmden Maurach	Multiple specimens. Fossilized rafts of +10 m long with complete colonies.	The Largest Know Crinoidean, from the family Pentacrinitidae. Among the tallest animals of its period, Seirocrinus is also one of the most famous fossils from the Posidonia Shale. It consists of fossils of colonies along large wood trunks, with specimens up to 14 m long, with the largest specimen reaching 26 m long,[188] what makes it among the tallest know Mesozoic organisms, one of the largest invertebrates know on the fossil record and one of the tallest know animals. It was an open ocean organism that lived in rafting woods, probably filtering food and serving as a refuge for other animals, such as ammonites.[189] The crinoids had a large colonization process, based on the status of the fossil wood found.[190][191] The large rafts where the home for a high variety of marine organisms, such as Balanoideans, Ammonites and other. It has been stimated that without the presence of modern raft wood predators (that apperared on the Bathonian) those rafts can last up to 5 years, being that the main reason the crinoids where able to attach such huge sizes. The large rafts where also proabaly essential to distribute animals along the Early Jurassic Seas.[192]	Close view of one specimen Single specimen
Pentacrinites[193]	Pentacrinites fossilis Pentacrinites briareus Pentacrinites dichotomus Pentacrinites quenstedti[194]	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Aselfingen Maurach Hemmikon Bascharage	Multiple specimens. Fossilized rafts of +10 m long with complete colonies.	Type genus of Crinoidean from the family Pentacrinitidae. Like Seirocrinus, Pentacrinites formed colonies on rafting wood, getting a different role than bigger crinoid and appearing on the first stages of the descomposition of the rafting wood. Was a smaller genus, with specimens of no more than 1 meter long, usually measuring 40–70 cm.	Close view of one specimen Reconstruction

Vertebrata

Fishes

Chondrichthyes

Genus	Species	Location	Material	Notes	Images
Microtoxodus[195]	Microtoxodus gulakmani[195]	Schömberg	Teeth	A non classified Elasmobranch.
Synechodus[196]	Synechodus egertoni	Holzmaden Ohmden Dotternhausen Schömberg Dudelange-Zoufftgen	Teeth	Thype shark of the family Synechodontiformes.
Rhomphaiodon[195]	Rhomphaiodon sp.	Schömberg Dudelange-Zoufftgen	Teeth	A shark of the family Synechodontiformes.
Protospinax[195][104]	Protospinax sp.	Schömberg Dudelange-Zoufftgen	Teeth	A member of the family Protospinacidae.
Palidiplospinax[197]	Palidiplospinax smithwoodwardi	Holzmaden	Articulated vertebral column, girdles, both fin spines and clasper organ	A member of the family Palaeospinacidae.
Palaeospinax[198]	Palaeospinax priscus	Holzmaden	Anterior part of body with basicranium, palatoquadrates, Meckel's cartilage, ceratohyals, epihyals, teeth, traces of the branchial arches and the anterior finspine	Type member of the family Palaeospinacidae.
Mesiteia[195]	? Mesiteia sp.	Schömberg Dudelange-Zoufftgen	Teeth	A Carpet shark of the family Orectolobiformes.
Chiloscyllium[195]	? Chiloscyllium sp.	Schömberg Dudelange-Zoufftgen	Teeth	A Carpet shark of the family Hemiscylliidae.	Modern Chiloscyllium
Palaeobrachaelurus[195][104]	Palaeobrachaelurus sp.	Schömberg Dudelange-Zoufftgen	Teeth	A Carpet shark of the family Orectolobiformes.
Ornatoscyllium[104]	Ornatoscyllium sp.	Dudelange-Zoufftgen	Teeth	A Carpet shark of the family Orectolobiformes.
Annea[104][199]	Annea sp.	Dudelange-Zoufftgen	Teeth	A Carpet shark of the family Orectolobiformes.
Paracestracion[104]	Paracestracion sp.	Dudelange-Zoufftgen	Teeth	A Bullhead Shark of the family Heterodontidae.
"Heterodontus"[104]	"Heterodontus" duffini	Dudelange-Zoufftgen	Teeth.	A Bullhead Shark of the family Heterodontidae.
Agaleus[195]	? Agaleus sp.	Dudelange-Zoufftgen	Teeth	A shark of the family Agaleidae.
Asteracanthus[195]	Asteracanthus sp.	Dudelange Holzmaden Ohmden Dotternhausen	Teeth	A shark of the family Acrodontidae.
Lissodus[195]	Lissodus guenneguesi Lissodus sp.	Dudelange-Zoufftgen	Teeth	A shark of the family Hybodontiformes.
Crassodus[200]	Crassodus reifi	Dotternhausen	Meckelian Cartilages, Jaws, teeth, Palatoquadrates, placoid scales and desarticualted parts of the labial, hyoid and branchial skeleton.	A shark of the family Hybodontidae. The Type specimen belongs to a large hybodontid, with an stimated total length of up to 3 m.[200] It has a Meckelian Cartilage more robust than Hybodus hauffianus and an acrodontine dentition, as seen on Bdellodus.[200] Probably is related with "Hybodus" delabechei, very likely a junior synonym of Crassodus.[200]
Bdellodus[201]	Bdellodus bollensis	Holzmaden Ohmden Dotternhausen	Teeth.	A shark of the family Hybodontidae.	Bdellodus
Hybodus[202][203]	Hybodus hauffianus	Banz Holzmaden Ohmden Dotternhausen Altdorf	Complete Specimens and fragments, including Teeth, scales, jaws and Cartilage.	Type shark of the family Hybodontidae. It is the most abundant shark on the layers of the Posidonia Shale, with some of the best preserved specimens of the genus known. It was probably an open ocean hunter, with small horns over the eyes. With a size around 2 m, it was also one of the largest representatives of the Chondrichthyes on the formation.[204][205]	Hybodus Holzmaden specimen, among the best preserved of the genus, with Belemmnites inside. Hybodus Another Holzmaden specimen
Pseudonotidanus[206]	Pseudonotidanus politus	Holzmaden Ohmden Dotternhausen	Fragmentary and Complete Specimens	A shark of the family Hexanchiformes.
Bathytheristes[207]	Bathytheristes gracilis	Ohmden	Partially Complete Specimen, with associated teeth	A member of Callorhynchidae inside Chimaeriformes. Similar to Callorhinchus, among the oldest known of its type.	Callorhinchus milii can be the closest relative of Bathytheristes
Acanthorhina[208]	Acanthorhina jaekeli	Holzmaden	Head and postcranial remains	A member of Myriacanthidae inside Chimaeriformes. An aberrant Chimaera with an extrange elongated nose and horns over the skull.	Acanthorhina
Metopacanthus[209]	Metopacanthus bollensis	Holzmaden Ohmden	Head and several postcranial remains	A member of Myriacanthidae inside Chimaeriformes. An aberrant Chimaera with a second jaw-like structure on its head.
Toarcibatis[104][210]	Toarcibatis elongata Toarcibatis sp.	Dudelange-Zoufftgen Longlaville	Teeth	A Basal member of the Rajiformes of the family Archaeobatidae.
Cristabatis[104][210]	Cristabatis crescentiformis Cristabatis sp.	Dudelange-Zoufftgen Longlaville	Teeth	A Basal member of the Rajiformes of the family Archaeobatidae.
Doliobatis[210]	Doliobatis weisi	Dudelange-Zoufftgen	Teeth	A Basal member of the Rajiformes of the family Archaeobatidae.
Jurobatos[104][199]	Jurobatos cappettai Jurobatos sp.	Dudelange-Zoufftgen	Teeth	A Basal member of the Rajiformes.

Actinopteri

Genus	Species	Location	Material	Notes	Images
Holzmadenfuro[211]	Holzmadenfuro rebmanni	Holzmaden	Complete Specimen	First ganoin-scaled Ophiopsiformes (Halecomorphi) from the Posidonienschiefer. The type specimen measures 51 cm, and has elongated and serrated body scales before the dorsal fin and tiny ganoid scales after it.[211]
Ohmdenfuro[211]	Ohmdenfuro bodmani	Ohmden	Nearly complete specimen with broken skull	First ganoin-scaled Ophiopsiformes (Halecomorphi) from the Posidonienschiefer. Elongated morphology, with a length of ~39 cm, covered by smooth, masive ganoin scales.[211]
Caturus[212]	Caturus smithwoodwardi	Holzmaden Würtenmberg	Various Complete and nearly complete Specimens	Type Genus of the family Caturidae inside Amiiformes	Caturus
Saurorhynchus[213]	Saurorhynchus hauffi Saurorhynchus brevirostris	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Maurach	Multiple specimens.	The youngest representative of the family Saurichthyidae, known for its large jaws, similar to modern Belonidae.	Saurorhynchus
Ptycholepis[214][215]	Ptycholepis bollensis Ptycholepis barrati	Dotternhausen Holzmaden Ohmden	Multiple specimens.	Type member of the family Ptycholepididae inside Ptycholepiformes. It is one of the Youngest representatives of its Family.	Ptycholepis Ptycholepis
Pholidophorus	Pholidophorus germanicus Pholidophorus hartmanni	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Maurach	Multiple specimens.	A member of the family Pholidophoridae. Is among the most abundant fishes on the late liassic of Europe, present on the sub-mediterranean boreal, with specimens of several sizes.	Pholidophorus Pholidophorus
Luxembourgichthys[216]	Luxembourgichthys friedeni	Bommelscheier industrial area, Bascharage	Multiple specimens.	A member of the family Pholidophoridae.
Leptolepis[217]	Leptolepis jaegeri Leptolepis coryphaenoides Leptolepis bronni Leptolepis normandica	Banz Altdorf Oedhof Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Maurach	Multiple specimens.	A member of the family Leptolepididae. The most common member of its family, Leptolepis is commonly associated with Crustaceans and small marine invertebrates, probably main creatures on its diet. One on the most predated vertebrates on the formation, with abundance of larger fishes and reptiles with specimens associated.	Leptolepis Leptolepis
Paraleptolepis[218]	Paraleptolepis wiedenrothi	SW of Braunschweig	MB. f.7612, nearly complete specimen.	A member of the family Leptolepididae. It differs from Leptolepis coryphaenoides in the presence of a few autapomorphics and also in the retention of several primitive features not present on the last one.[218] Small genus, of about 14 cm length.[218]
Euthynotus[219]	Euthynotus incognitus	Holzmaden Ohmden	Multiple specimens.	A member of the family Pachycormidae.	Euthynotus
Saurostomus[220]	Saurostomus esocinus	Banz Altdorf Dotternhausen Holzmaden Ohmden	Multiple specimens.	A member of the family Pachycormidae. Large representative of the family, reaching sizes up to 2.3 m.	Saurostomus Saurostomus
Pachycormus[221][222]	Pachycormus macropterus Pachycormus bollensis	Banz Altdorf Dotternhausen Holzmaden Ohmden	Multiple specimens.	Type member of the family Pachycormidae. Large representative of its family, with a size up to 1.5 m. One specimen preserved the alimentary canal, with the stomach filled by numerous hooklets that can be referred to the coleoid cephalopod Phragmoteuthis, impliying a diet of cephalopods from this genus.[223]	Pachycormus
Sauropsis[224][225]	Sauropsis latus Sauropsis veruinalis[212]	Holzmaden Ohmden	Multiple specimens.	A large member of the family Pachycormidae.	Sauropsis Sauropsis
Ohmdenia[226]	Ohmdenia multidentata	Ohmden	Partial specimen.	A large member of the family Pachycormidae, with up to 2.5–3 m long and an estimated weight over 200 kg.[226][227] Considered originally a junior synonym of Pachycormus, altrougth the craneal bones suggest a new genus. Among the largest fish found on the formation it is a key fossil on the transition to large filter feeding fishes.[227] Being Coeval with another basal Pachycormiformes show the specialization of the group during the late lower jurassic.[227] Ohmdenia is the sister taxa to group of suspension-feeding Giant Middle-Late Jurassic Fishes (Including the famous Leedsichthys), showing alterations on its dental structure, with jaw indicates a diet based on soft body prey.[227] Its evolutionary significance is equiparable to that of the genus Aetiocetus for the modern Baleen whale.[227]	Ohmdenia
Dapedium[228]	Dapedium pholidotum[229] Dapedium punctatus Dapedium caelatus Dapedium stollorum[230]	Banz Altdorf Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden Maurach	Multiple specimens.	A deep-bodied neopterygian, Type member of the family Dapediidae. Unpublished material indicates the presence of one or even two more still undescribed species of Dapedium in the Lower Toarcian.[229]	Dapedium Dapedium
Lepidotes[231]	Lepidotes elvensis Lepidotes gigas	Banz Mistelgau Aichelberg Schandelah Niedersachsen Dotternhausen Holzmaden Ohmden	Multiple specimens.	A common member of the Lepisosteiformes.	Lepidotes Lepidotes
Tetragonolepis[232]	Tetragonolepis drosera Tetragonolepis semicincta	Banz Altdorf Mistelgau Aichelberg Schandelah Dotternhausen Holzmaden Ohmden Maurach	Multiple specimens.	A common member of the Semionotidae.	Tetragonolepis
Strongylosteus[233][234]	Strongylosteus hindenburgi	Holzmaden Ohmden Dotternhausen	Various Specimens	A large member of the Chondrosteidae and the largest non reptilian marine vertebrate of the Posidonia shale, with a size between 3 and 4.5 m, and an estimated weight over 800 kg to 1 tonne.[233] Has been suggested as a junior Synonym of Chrondrosteus, altrougth there haven't been any new revision about the status of the genus. It is related to modern sturgeons, but with a different kind of mouth than common species, made for hunting prey in open waters, with a strong lower jaw, similar to modern Beluga Sturgeons.[235]	Strongylosteus

Sarcopterygii

Genus	Species	Location	Material	Notes	Images
Trachymetopon[236][237]	Trachymetopon liasicum	Holzmaden Ohmden Dotternhausen	Various Specimens with diverse degree of preservation.	A large coelacanth of the family Mawsoniidae, related with the genera Axelrodichthys, Chinlea, Diplurus and the type Mawsonia.[237] The largest specimen know from the Posidonia Shale is GPIT.OS.770 (Holotype), with a length over 1.6 m.[236] The specimen presents an ossified lung inside the abdominal cavity, and most of the body, being also one of the most complete Coelacanths of the Jurassic found.[236][237] Some recent discoveries from the Middle Jurassic show specimens of up to 3.5 m long.[238] Trachymetopon precedes the presence of the family Mawsoniidae in Europe by about 120 Ma and the northernmost occurrence of a member of the group, impliying an extensive geographical range during the Early Jurassic.[237] Due to the specimens being found on pelagic deposits suggest that probably was an open ocean swimmer.[236][237]	Trachymetopon GPIT.OS.770

Reptiles

Ichthyosauria

Inderminate specimens are known.[99]

Genus	Species	Location	Material	Notes	Images
Temnodontosaurus[239][240]	Temnodontosaurus trigonodon Temnodontosaurus burgundiae Temnodontosaurus "sp. A" Temnodontosaurus "sp. B"	Banz Altdorf Mistelgau Schlierbach Niedersachsen Holzmaden Ohmden Dotternhausen	Multiple specimens with different degree of preservation.	Type genus of the family Temnodontosauridae. A large Macroraptorial Ichthyosaur, apex predator of its environment. It range on sizes between the 9 and the 12 m, being one of the largest know ichthyosaurs, characterized by skulls and jaws over 1 m in length, with the largest being over 1.9 m long. It has been found with fragments of young icthyosaur on his stomach.[241] Of 39 specimens of Temnodontosaurus studied, a 21% (8 Specimens) show pathologies along their body, with several injuries post traumatic, probably done by other marine reptiles.[242]	Temnodontosaurus hunting Temnodontosaurus trigonodon specimen.
Stenopterygius[243][244][245][246]	Stenopterygius megalorhinus[243][244][245][246] Stenopterygius eos[243][244][245][246] Stenopterygius intercedens[243][244][245][246] Stenopterygius incessus[243][244][245][246] Stenopterygius hauffianus[243][244][245][246] Stenopterygius cuneiceps[243][244][245][246] Stenopterygius promegacephalus[243][244][245][246] Stenopterygius sp[243][244][245][246]	Banz Altdorf Mistelgau Schlierbach Niedersachsen Holzmaden Ohmden Dotternhausen	Multiple specimens. Some of them are among the best preserved Ichthyosaur remains known.[243][244][245][246]	Type genus of the family Stenopterygiidae. A common Toarcian Ichthyosaur, present on multiple layers. The rather exquisite level of preservation has led to know even the coloration, that exposes a clear countershading, with an upper part being more obscure than the lower, similar to modern Killer Whales, the Heaviside's dolphin or the Dall's porpoise. There is also evidence of changes in color with ontogenic chnges, going from dark juveniles to countershaded adults. The skin was flexible & Scaless, as in Dolphins.[247]	Restoration Fossil mother with embryo
Suevoleviathan[248][249]	Suevoleviathan disinteger Suevoleviathan integer[250]	Holzmaden Ohmden Banz Dotternhausen	Multiple specimens.	Type genus of the family Suevoleviathanidae. Includes specimens up to 4 m long.	Suevoleviathan integer fossil
Hauffiopteryx[251]	Hauffiopteryx typicus	Holzmaden Ohmden Dotternhausen	Complete Specimen.	Small sized Ichthyosaur, probably a member of Parvipelvia.[252]	Hauffiopteryx typicus fossil
Eurhinosaurus[253][254][255]	Eurhinosaurus longirostris	Banz Altdorf Holzmaden Ohmden Dotternhausen	Complete and partial Specimens.	A large ichthyosaur of the family Leptonectidae with convergent evolution with modern Swordfish. Like this fishes, Eurhinosaurus is believed to be a fast swimming predator, able to hunt fish schools on same way. Large specimens of up to 6 m are known.	Complete specimen from the Posidonia Shale Eurhinosaurus restoration

Plesiosauria

Genus	Species	Location	Material	Notes	Images
Meyerasaurus[256]	Meyerasaurus victor	Dotternhausen Holzmaden	Nearly complete specimen.	A Rhomaleosauridae Plesiosaur. Its detailed fossils have helped to study plesiosaur movement.[257]	Meyerasaurus
Hauffiosaurus[258]	Hauffiosaurus zanoni	Dotternhausen Holzmaden	Complete specimen.	A Rhomaleosauridae Plesiosaur. A moderately sized (3.4 m) Rhomaleaosaurid, ecologically adapted to fish hunt, as has been observed due to comparing the long snouted skull with that of Peloneustes, Gharial Crocodiles or Dolphins.[258]	Hauffiosaurus
Plesiopterys[259][183]	Plesiopterys wildi[260]	Holzmaden	Complete Specimen.	A basal Plesiosaur that has been linked with Cryptoclididae. It is one of the smallest from the Posidonia, with a complete skeleton measuring less than 2.5 m. It is considered a possible junior synonym of Seeleyosaurus[261]	Plesiopterys
Seeleyosaurus[262][263]	Seeleyosaurus guilelmiimperatoris	Holzmaden	Complete Specimens.	A Plesiosaur of the family Microcleididae. It was named originally "Plesiosaurus guilelmiimperatoris". It was a moderate‐sized plesiosauroid, measuring up to 3,5 m in length with a skull length of 170 mm.[263]	Seeleysaurus
Microcleidus[264]	Microcleidus brachypterygius	Holzmaden	Several Complete Specimens.	Type member of the Plesiosaur family Microcleididae. Small Plesiosaur, with a length of less than 3 m. Possible Junior synonym of Hydrorion	Microcleidus
Hydrorion[265][263][266]	Hydrorion brachypterygius	Holzmaden	Complete Specimen.	A Plesiosaur of the family Plesiosauridae. It is characterised by a really enlongated neck, was probably an ichthyophagous form that occurred rarely in the Posidonienschiefer fauna.	Hydrorion

Sphenodontia

Genus	Species	Location	Material	Notes	Images
Palaeopleurosaurus[267]	Palaeopleurosaurus posidoniae	Banz Holzmaden Ohmden Dotternhausen Kerkhofen	Complete specimens.	An aquatic sphenodont of the family Pleurosauridae. Palaeopleurosaurus evidences that there was a slightly skeletal specialization for an aquatic lifestyle, achieved trougth the Jurassic gradually on pleurosaurs.[267] It has similarities with other marine reptiles, such with members of Sauropterygia the presence of a defined suture between the centrum and the neural arch, along with reducted sternum.[268] Probably had a semiaquatic style of life, altrought not as adapted as Pleurosaurus, as show limited morphological evidence of adaptation to a complete aquatic lifestyle, defined by no Osteosclerosis and the lack of Pachyostosis, except for a thicker shaft region in the humerus, that is as narrow as in terrestrial rhynchocephalians, such as the terrestrial Clevosaurus.[268] Palaeopleurosaurus probably was still able to walk on land, for example for Oviposition.[268] Recent studies suggest a shorter lifespan than modern Tuatara, based on irregular spacing of growth marks.[269]	Palaeopleurosaurus Palaeopleurosaurus

Testudinata

Genus	Species	Location	Material	Notes
Testudinata[270][271][272]	Testudinata indeterminate A Testudinata indeterminate B Testudinata indeterminate C	Altdorf, Bavaria	Various partial specimens, mostly pleurals.	Unclassified Testudine. Münster (1834) cited: " there were also rare things at the quarries of Altdorf, among other remains there where ones of a turtle on lias limestone"". The remains are not cataloged.[273]
Mesochelydia?[274]	Mesochelydia? indeterminate A Mesochelydia? indeterminate B	Banz, Bavaria	Various partial specimens, mostly pleurals. Includes a left partial shell.	Assigned to Mesochelydia(?) due to resemblances for the authors to modern turtles. Unclassified and non cataloged.[273]
Perichelydia?[275]	Perichelydia? indeterminate A	Schandelah	Fragmentary Remains	Can be a more basal taxon or even not a turtle. If it is, can be the first Turtle remains found on the Posidonia Shale since the late 1800s. Now on a private collection.[275]

Crocodrylomorpha

Genus	Species	Location	Material	Notes	Images
Steneosaurus[276][102]	Steneosaurus bollensis	Banz Altdorf Mistelgau Schlierbach Niedersachsen Holzmaden Ohmden Dotternhausen	Complete specimens.	A marine crocodrylomorph with a diet probably based on fish.[277] It is the best known member of the family Teleosauriade. The Genus was declared Invalid in 2020.
Macrospondylus[278]	Macrospondylus bollensis	Banz Altdorf Mistelgau Schlierbach Niedersachsen Holzmaden Ohmden Dotternhausen	Series of Complete specimens, including skin impressions	A Thalattosuchian of the family Teleosauridae. Was considered synonymous with Steneosaurus until in 2020 this last was recovered as invalid.[279] Includes 5 m specimens.	Macrospondylus bollensis fossil with Skin Impressions
Mystriosaurus[280][281]	Mystriosaurus laurillardi	Banz Altdorf Holzmaden Ohmden Dotternhausen	Complete specimen.	A Thalattosuchian of the family Teleosauridae. A marine crocodrylomorph with a diet probably based on fish. Was considered synonymous with Steneosaurus until recently.[282]	Mystriosaurus
Platysuchus[283]	Platysuchus multiscrobiculatus	Banz Altdorf Schandelah Niedersachsen Holzmaden Ohmden Dotternhausen	Complete specimen.	A Thalattosuchian of the family Teleosauridae. Platysuchus was slightly more robust than its contemporaneous relatives, being probably adapted to hunt more voluminous fish.	Platysuchus
Pelagosaurus[284][285]	Pelagosaurus typus	Banz Altdorf Ohmden Holzmaden Dotternhausen	Complete specimen.	A Thalattosuchian with a complex assignation, where can be a member of the family Teleosauridae or the basalmost Metriorhynchoidean. With a less heavy built, bigger skull and more visible marine adaptation, its diet probably based completely on fish (one fossil specimen was found with a Leptolepis in its stomach contents). Has been considered a basal Metriorhynchidae[286] or sister taxon to both Teleosauridae and Metriorhynchidae.[287]	Pelagosaurus
Pholidosauridae?[276]	Pholidosauridae Indeterminate	Holzheim bei Göppingen	Partial specimen.	A possible Pholidosauridae Neosuchian. A marine crocodrylomorph, that can be a mistaken Teleosaurian as has been included on Cf. Platysuchus, altrougth has characters that align it with Anglosuchus, that was also confused with a Teleosaurid.[276]
Goniopholididae?[102]	Goniopholididae Indeterminate	Altdorf	Frontal Snout	A Possible Goniopholid Neosuchian. Classified as "?Steneosaurus" sp., it was noted initially a morphology more similar to Goniopholis simus and specially Amphicotylus lucasii, being a possible early member of the group.[102]
Protosuchidae?[275]	Protosuchidae Indeterminate	Schandelah	Fragmentary remains	Possible member of the Protosuchia inside Crocodyliformes. If is confirmed as a member of the family it would be a terrestrial, maybe semi-acuatic crocodrylomorph. Can be a mistaken juvenile Teleosaurian.[275]

Pterosauria

Genus	Species	Location	Material	Notes	Images
Campylognathoides[288][289][290]	Campylognathoides zitteli[290] Campylognathoides liasicus[290]	Banz Altdorf Niedersachsen Holzmaden Ohmden Dotternhausen	Several specimens with different degrees of preservation.	A Novialoidean Pterosaur, type genus of the family Campylognathoidea. Mark Witton suggests the construction of Campylognathoides' extremely robust forelimbs, with proportionally long wing fingers, could be a specialization for a fast aerial lifestyle comparable to those of Falcons and mastiff bats, being more probably an insect & vertebrate hunter and living on nearshore environments.[291]	Campylognathoides Nearly complete Campylognathoides
"Schandelopterus"[292]	Novialoidea ("Schandelopterus") Indeterminate	Schandelah	Pelvis and several vertebrae.	A Novialoidean Pterosaur, probably a member of the family Campylognathoidea. Has been assigned to the genus Campylognathoides, although it is clearly different than any other pterosaur from the Posidonia Shale.[292] The name "Schandelopterus" is invalid and lacks any study, assigned without species to refer to the specimen on private German Fossil Groups. The pelvis indicates a laterally, slightly upwardly directed orientation of the acetabula which does not support a bird-like bipedal locomotion of this pterosaur as has been suggested on the past.[292]
Parapsicephalus[293]	cf. Parapsicephalus purdoni	Altdorf	Skull	A Rhamphorhynchinae Pterosaur. Has been assigned to the genus Dorygnathus. It has a really complete skull that can help to explain the estatus of the genus Parapsicephalus.[293]
Dorygnathus[294][295]	Dorygnathus banthensis[296] Dorygnathus mistelgauensis?[297]	Banz Altdorf Schandelah Niedersachsen Holzmaden Ohmden Dotternhausen	Multiple specimens with different degree of preservation.[298]	A Rhamphorhynchinae Pterosaur. It is one of the best known Early Jurassic Pterosaurs.[294] Unlike Campylognthoides, Dorygnathus was an oceanic hunter, with teeth disposed to catch marine prey, such as Belemnittes and several species of fishes. Dorygnathus mistelgauensis is considered a junior synonym until more data can be recovered from the specimen, held on a private collection.[294]	Dorygnathus on terrestrial pose Nearly complete Dorygnathus

Dinosauria

Genus	Species	Location	Material	Notes	Images
Neotheropoda[299]	Neotheropoda? Indeterminate	Schandelah	Cervical Vertebrae, dorsal vertebrae, limbs and possible ulna	An indeterminate possible Theropod dinosaur, possibly Neotheropoda. Hasn't been revised since 1984.
Sauropodiformes[300]	Sauropodiformes Indeterminate	Oedhof	3 cm long, tooth-studded fragment of a lower jaw	An indeterminate possible Sauropodomorph dinosaur, possibly a member of Sauropodiformes (Resemble Yunnanosaurus teeth).[300] The Fossil was reported from the Lias Epsilon level, that on Oedhof is occupied by the Posidonia Shale.[300] It was found with abundant Plant debris and Belemnite remains.[300] Hasn't been revised since 1956
Ohmdenosaurus[301]	Ohmdenosaurus liasicus	Ohmden	Tibia and astragalus	A Gravisaurian Sauropod. One of the few formally described from the Toarcian. Has been related with Vulcanodon, although more recent studies placed it as a relative of Rhoetosaurus.[302] It has been claimed to be a small sauropod with a size of 4 m, altrought the tibia measures 405 to 410 mm, leading to a 6.7 m long sauropod. Molina Pérez & Larramendi, stimated a modern size of 6.2 to 6.7 m long, with a weight of 1.3 tonnes.[303]	Ohmdenosaurus

Synapsida

Genus	Species	Location	Material	Notes	Images
Cynodontia?[304]	Cynodontia? Indeterminate	Banz	3 Teeth	The specimens were listed on several notes on the 1800s (As "mammal teeth"), although its existence has not been proved. Can be related to the Trithelodontidae or other late surviving Cynodonts by related characters, such as Irajatherium, but also to the Mammaliformes. If its presence is proved, it would be the first Synapsid found on the Posidonia Shale.

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