Orchideae

Orchideae is a tribe of orchids in the subfamily Orchidoideae. It is the largest tribe, containing more than 1,700 species. It has been divided into two subtribes, Orchidinae and Habenariinae. However, although phylogenetic studies have established the monophyly of the subtribes, the generic boundaries are unclear, with many genera as traditionally circumscribed being paraphyletic or even polyphyletic.[1] Species of genera such as Habenaria and Platanthera are placed into both subtribes,[2] so a clear assignment of all genera to subtribes is not possible as of March 2018.

Orchideae
Flower of Orchis provincialis
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Monocots
Order:	Asparagales
Family:	Orchidaceae
Subfamily:	Orchidoideae
Tribe:	Orchideae (Dressler & Dodson) Verm. (1977)
Subtribes
Orchidinae Habenariinae

The Asian species of Orchideae, in particular, have been subject to repeated changes of generic placement from 2012 onwards.[2][1][3][4][5]

Prairie White Fringed Orchid (Platanthera leucophaea)

Genera include:

• Anacamptis, Bartholina, Benthamia, Bonatea, Brachycorythis, Centrostigma, Chamorchis, Cynorkis, Dactylorhiza, Diplomeris, Dracomonticola, Galearis, Gennaria, Gymnadenia, Habenaria, Hemipilia, Herminium, Himantoglossum, Holothrix, Megalorchis, Neobolusia, Neotinea, Oligophyton, Ophrys, Orchis, Pecteilis, Peristylus, Physoceras, Platanthera, Platycoryne, Ponerorchis, Pseudorchis, Roeperocharis, Schizochilus, Serapias, Sirindhornia, Stenoglottis, Steveniella, Thulinia, Traunsteinera, Tsaiorchis, Tylostigma, Veyretella

Genera included in a 2014 molecular phylogenetic study are listed below, together with subtribe clades into which their species were placed. Four genera were split between the two subtribes.[1]

• Bonatea – Habenariinae
• Dactylorhiza – Orchidinae
• Galearis (including Neolindleya) – Orchidinae
• Gennaria – Habenariinae
• Gymnadenia – Orchidinae
• Habenaria – Habenariinae
• Hemipilia (including Hemipiliopsis) – Orchidinae
• Herminium (including Androcorys, Porolabium) – Habenariinae, Orchidinae
• Ophrys – Orchidinae
• Orchis – Orchidinae
• Pecteilis – Habenariinae
• Peristylus – Habenariinae, Orchidinae
• Platanthera (including Smithorchis) – Habenariinae, Orchidinae
• Ponerorchis (including Amitostigma, Neottianthe) – Habenariinae, Orchidinae
• Pseudorchis – Orchidinae
• Serapias – Orchidinae
• Sirindhornia – Orchidinae
• Tsaiorchis – Orchidinae