Nepenthes angasanensis

Nepenthes angasanensis
	Nepenthes angasanensis holotype (Salmon & Maulder 234372).
Conservation status
	; Least Concern (IUCN 3.1)[1]
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Order:	Caryophyllales
Family:	Nepenthaceae
Genus:	Nepenthes
Species:	N. angasanensis
Binomial name
	Nepenthes angasanensis; Maulder, D.Schub., B.R.Salmon & B.Quinn (1999)[2]
	Distribution of N. angasanensis.
Synonyms
	Synonyms Nepenthes mikei; auct. non B.R.Salmon & Maulder:; Jebb & Cheek (1997);[3]; Cheek & Jebb (2001)[4]; [=N. angasanensis/N. mikei] ; Nepenthes tobaica; auct. non Danser (1928): Danser (1940)[5] ; Nepenthes tobaica; auct. non Danser: Tamin & M.Hotta in M.Hotta (1986)[6]; [=N. angasanensis/N. tobaica] ;

Nepenthes angasanensis /nɪˈpɛnθiːz ˌæŋɡəsəˈnɛnsɪs/ is a tropical pitcher plant species endemic to Sumatra, where it grows at an altitude of 1,400 metres (4,600 ft) to 3,100 metres (10,200 ft) above sea level.[1][7] The status of this taxon is controversial as it is similar in morphology to N. mikei and N. tobaica. It has even been suggested that the taxon might represent a natural hybrid between N. densiflora and N. tobaica.[8]

The specific epithet refers to Mount Puncak Angasan, from which the type specimen was collected.[2] No forms or varieties of N. angasanensis have been described.

Taxonomy

Differences between *N. angasanensis*, *N. mikei* and *N. tobaica* (Salmon & Maulder, 1999)
Character	N. angasanensis	N. mikei	N. tobaica
Habit	Produces offshoots from underground rhizomes	No rhizomes	No rhizomes
Spur	Forked	Fasciculate	Filiform
Inner margin of peristome	Teeth to 1.5 millimetres (0.06 in) to 2 millimetres (0.08 in) long	Teeth to 0.2 millimetres (0.01 in) to 0.4 millimetres (0.02 in) long	Teeth < 0.2 millimetres (0.01 in)
Stem cross section	Cylindrical	Cylindrical	Cylindrical to obtusely triangular
Bracteoles	Sometimes near base of lowest pedicel only	Half way up every pedicel	At base or slightly below pedicel attachment, few
Pitcher glands	300 / cm² (1900 per sq in)	150-180 / cm² (1000 to 1200 per sq in)	200-250 / cm² (1300 to 1600 per sq in)
Pedicels	1-flowered	1-flowered	2-flowered
Inflorescence (female)	5.5 centimetres (2 in) to 12.5 centimetres (5 in) long, 9-17 flowers	4 centimetres (2 in) to 8 centimetres (3 in) long, 4-10 flowers	19.5 centimetres (8 in) to 40 centimetres (16 in) long, 30-50 flowers

In 2001, Charles Clarke performed a cladistic analysis of the Nepenthes species of Sumatra and Peninsular Malaysia using 70 morphological characteristics of each taxon. The following is a portion of the resultant cladogram, showing part of "Clade 6". The sister pair of N. angasanensis and N. mikei has 79% support.[9]

unnamed

	N. gracilis

	N. reinwardtiana

unnamed

N. tobaica

79%

	N. angasanensis

	N. mikei

Natural hybrids

The following natural hybrids involving N. angasanensis have been recorded.

N. angasanensis × N. densiflora[9]

References

Clarke, C.M. (2018). "Nepenthes angasanensis". IUCN Red List of Threatened Species. 2018: e.T39642A143958899. Retrieved 18 April 2020.
Salmon, B. & R. Maulder 1999. Notes on Nepenthes from Northern Sumatra. Carnivorous Plant Newsletter 28(1): 14–18.
Jebb, M.H.P. & M.R. Cheek 1997. A skeletal revision of Nepenthes (Nepenthaceae). Blumea 42(1): 1–106.
Cheek, M.R. & M.H.P. Jebb 2001. Nepenthaceae. Flora Malesiana 15: 1–157.
Danser, B.H. 1940. A new Nepenthes from Sumatra. Bulletin du Jardin Botanique de Buitenzorg, Série III, 16: 268–271.
(in Indonesian) Tamin, R. & M. Hotta 1986. Nepenthes di Sumatera: The genus Nepenthes of the Sumatra Island. In: M. Hotta (ed.) Diversity and Dynamics of Plant Life in Sumatra: Forest Ecosystem and Speciation in Wet Tropical Environments. Part 1: Reports and Collection of Papers. Kyoto University, Kyoto. pp. 75–109.
McPherson, S.R. & A. Robinson 2012. Field Guide to the Pitcher Plants of Sumatra and Java. Redfern Natural History Productions, Poole.
Schlauer, J. N.d. Nepenthes angasanensis. Carnivorous Plant Database.
Clarke, C.M. 2001. Nepenthes of Sumatra and Peninsular Malaysia. Natural History Publications (Borneo), Kota Kinabalu.

Clarke; et al. (2000). "Nepenthes angasanensis". IUCN Red List of Threatened Species. 2000. Retrieved 11 May 2006.CS1 maint: ref=harv (link)old-form url
Hernawati & P. Akhriadi 2006. A Field Guide to the Nepenthes of Sumatra. PILI-NGO Movement, Bogor.

External links

Photographs of N. angasanensis at the Carnivorous Plant Photofinder

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[IUCN-1] Clarke, C.M. (2018). "Nepenthes angasanensis". IUCN Red List of Threatened Species. 2018: e.T39642A143958899. Retrieved 18 April 2020.

[angasan-2] Salmon, B. & R. Maulder 1999. Notes on Nepenthes from Northern Sumatra. Carnivorous Plant Newsletter 28(1): 14–18.

[J&C-3] Jebb, M.H.P. & M.R. Cheek 1997. A skeletal revision of Nepenthes (Nepenthaceae). Blumea 42(1): 1–106.

[C&J-4] Cheek, M.R. & M.H.P. Jebb 2001. Nepenthaceae. Flora Malesiana 15: 1–157.

[5] Danser, B.H. 1940. A new Nepenthes from Sumatra. Bulletin du Jardin Botanique de Buitenzorg, Série III, 16: 268–271.

[T&H-6] (in Indonesian) Tamin, R. & M. Hotta 1986. Nepenthes di Sumatera: The genus Nepenthes of the Sumatra Island. In: M. Hotta (ed.) Diversity and Dynamics of Plant Life in Sumatra: Forest Ecosystem and Speciation in Wet Tropical Environments. Part 1: Reports and Collection of Papers. Kyoto University, Kyoto. pp. 75–109.

[7] McPherson, S.R. & A. Robinson 2012. Field Guide to the Pitcher Plants of Sumatra and Java. Redfern Natural History Productions, Poole.

[Schlauer-8] Schlauer, J. N.d. Nepenthes angasanensis. Carnivorous Plant Database.

[Clarke-9] Clarke, C.M. 2001. Nepenthes of Sumatra and Peninsular Malaysia. Natural History Publications (Borneo), Kota Kinabalu.