Evolutionarily stable state

"A population is said to be in an evolutionarily stable state if its genetic composition is restored by selection after a disturbance, provided the disturbance is not too large. Such a population can be genetically monomorphic or polymorphic." —Maynard Smith (1982).

This contrasts with the definition of an evolutionarily stable strategy, of which is a game theory concept.

"An ESS or evolutionarily stable strategy is a strategy such that, if all the members of a population adopt it, no mutant strategy can invade." —Maynard Smith (1982).

Mathematical formulation

Evolutionarily stable states are frequently used to identify solutions to the replicator equation, given in its linear payoff form by:

{\dot {x_{i}}}=x_{i}\left(\left(Ax\right)_{i}-x^{T}Ax\right),

The state ${\hat {x}}$ is said to be evolutionarily stable if for all $x\neq {\hat {x}}$ in some neighborhood of ${\hat {x}}$ .

x^{T}Ax<{\hat {x}}^{T}Ax

This is similar but distinct from the concept of an evolutionarily stable strategy.

Properties

It can be shown that for the replicator equation if a state is evolutionarily stable then it is an asymptotically stable rest point, so the evolutionarily stable states are often taken as solutions of the replicator equation. The converse is true if the game matrix is symmetric.

The concept of evolutionary stability is equivalent to the concept of strong stability for normal form games, due to Cressman.

References

Maynard Smith, J.. (1982) Evolution and the Theory of Games. Cambridge University Press. ISBN 0-521-28884-3

Topics in game theory
Definitions	Cooperative game Determinacy Escalation of commitment Extensive-form game First-player and second-player win Game complexity Graphical game Hierarchy of beliefs Information set Normal-form game Preference Sequential game Simultaneous game Simultaneous action selection Solved game Succinct game
Equilibrium concepts	Nash equilibrium Subgame perfection Mertens-stable equilibrium Bayesian Nash equilibrium Perfect Bayesian equilibrium Trembling hand Proper equilibrium Epsilon-equilibrium Correlated equilibrium Sequential equilibrium Quasi-perfect equilibrium Evolutionarily stable strategy Risk dominance Core Shapley value Pareto efficiency Gibbs equilibrium Quantal response equilibrium Self-confirming equilibrium Strong Nash equilibrium Markov perfect equilibrium
Strategies	Dominant strategies Pure strategy Mixed strategy Strategy-stealing argument Tit for tat Grim trigger Collusion Backward induction Forward induction Markov strategy
Classes of games	Symmetric game Perfect information Repeated game Signaling game Screening game Cheap talk Zero-sum game Mechanism design Bargaining problem Stochastic game n-player game Large Poisson game Nontransitive game Global game Strictly determined game Potential game
Games	Go Chess Infinite chess Checkers Tic-tac-toe Prisoner's dilemma Gift-exchange game Optional prisoner's dilemma Traveler's dilemma Coordination game Chicken Centipede game Volunteer's dilemma Dollar auction Battle of the sexes Stag hunt Matching pennies Ultimatum game Rock paper scissors Pirate game Dictator game Public goods game Blotto game War of attrition El Farol Bar problem Fair division Fair cake-cutting Cournot game Deadlock Diner's dilemma Guess 2/3 of the average Kuhn poker Nash bargaining game Induction puzzles Trust game Princess and Monster game Rendezvous problem
Theorems	Arrow's impossibility theorem Aumann's agreement theorem Folk theorem Minimax theorem Nash's theorem Purification theorem Revelation principle Zermelo's theorem
Key figures	Albert W. Tucker Amos Tversky Antoine Augustin Cournot Ariel Rubinstein Claude Shannon Daniel Kahneman David K. Levine David M. Kreps Donald B. Gillies Drew Fudenberg Eric Maskin Harold W. Kuhn Herbert Simon Hervé Moulin Jean Tirole Jean-François Mertens Jennifer Tour Chayes John Harsanyi John Maynard Smith John Nash John von Neumann Kenneth Arrow Kenneth Binmore Leonid Hurwicz Lloyd Shapley Melvin Dresher Merrill M. Flood Olga Bondareva Oskar Morgenstern Paul Milgrom Peyton Young Reinhard Selten Robert Axelrod Robert Aumann Robert B. Wilson Roger Myerson Samuel Bowles Suzanne Scotchmer Thomas Schelling William Vickrey
See also	All-pay auction Alpha–beta pruning Bertrand paradox Bounded rationality Combinatorial game theory Confrontation analysis Coopetition Evolutionary game theory First-move advantage in chess Game mechanics Glossary of game theory List of game theorists List of games in game theory No-win situation Solving chess Topological game Tragedy of the commons Tyranny of small decisions

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