Pacullidae

Pacullidae
Scientific classification
Kingdom:Animalia
Phylum:Arthropoda
Subphylum:Chelicerata
Class:Arachnida
Order:Araneae
Infraorder:Araneomorphae
Family:Pacullidae
Simon, 1893[1]
Genera

See text.

Pacullidae is a family of araneomorph spiders. It was merged into Tetrablemmidae by Brignoli in 1973 and by Lehtinen in 1981.[1] In 2016, a large phylogenetic study proposed reviving the family.[2] As of May 2018, the family is accepted by the World Spider Catalog.[1]

Description

The family Pacullidae contains three-clawed spiders with six eyes, lacking a cribellum. They resemble spiders from the family Tetrablemmidae in some respects but are much larger, always exceeding 5 mm long, have a very wrinkled (rugose) cuticle, and females do not have large membranous receptacles.[2]

Phylogeny

Pacullidae falls within the Synspermiata clade, a clade of former haplogyne spiders with "synsperm" – encapsulated groups of 2–4 fused sperm cells. Within this clade, it groups with four other families, including Tetrablemmidae, but is distinct from the latter, being most closely related to Diguetidae. Together with Pholcidae, these four families are placed in the "lost trachea clade", a group of families that have lost their posterior respiratory system.[2]

"Lost trachea clade"

 

 

Tetrablemmidae

 
 

 

Plectreuridae

 
 

 

Diguetidae

 
 

Pacullidae

 
 
 
 
 

Pholcidae

 
 

Genera and species

As of May 2018, the World Spider Catalog accepts the following genera and species:[1]

  • Lamania Lehtinen, 1981
    • Lamania bernhardi (Deeleman-Reinhold, 1980) – Borneo
    • Lamania bokor Schwendinger & Košulič, 2015 – Cambodia
    • Lamania gracilis Schwendinger, 1989 – Bali
    • Lamania inornata (Deeleman-Reinhold, 1980) – Borneo
    • Lamania kraui (Shear, 1978) – Thailand, Malaysia
    • Lamania lipsae Dierkens, 2011 – Borneo
    • Lamania nirmala Lehtinen, 1981 (type species) – Borneo
    • Lamania sheari (Brignoli, 1980) – Indonesia (Sulawesi)
  • Paculla Simon, 1887
    • Paculla bukittimahensis Lin & Li, 2017 – Singapore
    • Paculla cameronensis Shear, 1978 – Malaysia
    • Paculla globosa Lin & Li, 2017 – Singapore
    • Paculla granulosa (Thorell, 1881) (type species) – New Guinea
    • Paculla mului Bourne, 1981 – Borneo
    • Paculla negara Shear, 1978 – Malaysia
    • Paculla sulaimani Lehtinen, 1981 – Malaysia
    • Paculla wanlessi Bourne, 1981 – Borneo
  • Perania Thorell, 1890
    • Perania annam Schwendinger & Košulič, 2015 – Vietnam
    • Perania armata (Thorell, 1890) – Indonesia (Sumatra)
    • Perania birmanica (Thorell, 1898) – Myanmar
    • Perania cerastes Schwendinger, 1994 – Malaysia
    • Perania coryne Schwendinger, 1994 – Malaysia
    • Perania deelemanae Schwendinger, 2013 – Indonesia (Sumatra)
    • Perania egregia Schwendinger, 2013 – Thailand
    • Perania ferox Schwendinger, 2013 – Thailand
    • Perania harau Schwendinger, 2013 – Indonesia (Sumatra)
    • Perania korinchica Hogg, 1919 – Indonesia (Sumatra)
    • Perania nasicornis Schwendinger, 1994 – Thailand
    • Perania nasuta Schwendinger, 1989 – Thailand
    • Perania nigra (Thorell, 1890) (type species) – Indonesia (Sumatra)
    • Perania picea (Thorell, 1890) – Indonesia (Sumatra)
    • Perania quadrifurcata Schwendinger, 2013 – Thailand
    • Perania robusta Schwendinger, 1989 – China, Thailand
    • Perania selatan Schwendinger, 2013 – Indonesia (Sumatra)
    • Perania siamensis Schwendinger, 1994 – Thailand
    • Perania tumida Schwendinger, 2013 – Thailand
    • Perania utara Schwendinger, 2013 – Indonesia (Sumatra)
  • Sabahya Deeleman-Reinhold, 1980
    • Sabahya bispinosa Deeleman-Reinhold, 1980 – Borneo
    • Sabahya kinabaluana Deeleman-Reinhold, 1980 (type species) – Borneo

References

  1. 1 2 3 4 "Family: Tetrablemmidae O. Pickard-Cambridge, 1873", World Spider Catalog, Natural History Museum Bern, retrieved 2018-05-19
  2. 1 2 3 Wheeler, Ward C.; Coddington, Jonathan A.; Crowley, Louise M.; Dimitrov, Dimitar; Goloboff, Pablo A.; Griswold, Charles E.; Hormiga, Gustavo; Prendini, Lorenzo; Ramírez, Martín J.; Sierwald, Petra; Almeida-Silva, Lina; Alvarez-Padilla, Fernando; Arnedo, Miquel A.; Benavides Silva, Ligia R.; Benjamin, Suresh P.; Bond, Jason E.; Grismado, Cristian J.; Hasan, Emile; Hedin, Marshal; Izquierdo, Matías A.; Labarque, Facundo M.; Ledford, Joel; Lopardo, Lara; Maddison, Wayne P.; Miller, Jeremy A.; Piacentini, Luis N.; Platnick, Norman I.; Polotow, Daniele; Silva-Dávila, Diana; Scharff, Nikolaj; Szűts, Tamás; Ubick, Darrell; Vink, Cor J.; Wood, Hannah M. & Zhang, Junxia (2016), "The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling", Cladistics, doi:10.1111/cla.12182
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