Flora of New York/Millerieae, Madieae, Eupatorieae

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Tribe Millerieae

Subtribe Milleriinae

Smallanthus

Smallanthus uvedalia

Hairy leafcup (Smallanthus uvedalia) was present in and collected from western New York during the first half of the twentieth century, but has not been confirmed since.

Asterales — Asteraceae — Asteroideae — Millerieae — Milleriinae — Smallanthus
Smallanthus	Smallanthus	N.Y. Status	Images	Distribution	NY NPT
Smallanthus uvedalia (L.) Mack. ex Small 1753. Osteospermum uvedalia L. 1763. Polymnia uvedalia (L.) L. 1933. Smallanthus uvedalius (L.) Mack. ex Small	Hairy leafcup, Bear's-foot, Yellow-flowered leafcup	Native, C:10, Endangered, No recent reports			NYFA-Z USDA-N0 NEW-0 ARS ITIS BNA Images, wsp

Guizotia

Niger-seed (Guizotia abyssinica) is commonly used to feed birds, which may account for its occasional temporary introduction.

Asterales — Asteraceae — Asteroideae — Millerieae — Milleriinae — Guizotia
Guizotia Cass.	Guizotia	N.Y. Status	Images	Distribution	NY NPT
Guizotia abyssinica (L.f.) Cass. 1782. Polymnia abyssinica L.f. 1829. Guizotia abyssinica (L.f.) Cass. 1836. Guizotia oleifera DC.	Niger-seed, Ramtilla	Introduced from Africa, Impersistent UPL Annual, Herb-forb			NYFA-X USDA-XW NPT Can ARS ITIS FNA Images, wsp

Subtribe Galinsoginae

Galinsoga

Galinsoga quadriradiata
shaggy soldier

Asterales — Asteraceae — Asteroideae — Millerieae — Galinsoginae
Galinsoga Ruiz & Pav.	Gallant-soldier	N.Y. Status	Images	Distribution	NY NPT
Galinsoga quadriradiata Cav. G. aristulata E.P.Bicknell G. bicolorata H.St.John & D.White G. caracasana (DC.) Sch.Bip. G. ciliata (Raf.) S.F.Blake	Fringed quickweed, Peruvian daisy, Shaggysoldier	Introduced			NYFA-X USDA-XX NPT Images, wsp
Galinsoga parviflora Cav. Galinsoga semicalva	Gallant-soldier, Small-flowered quickweed	Introduced			NYFA-X USDA-XX NPT Images, wsp

Subtribe Melampodiinae

Acanthospermum

Acanthospermum australe
Paraguay starbur

Asterales — Asteraceae — Asteroideae — Millerieae — Melampodiinae
Acanthospermum Schrank	Starbur	N.Y. Status	Images	Distribution	NY NPT
Acanthospermum australe (Loefl.) Kuntze 1758. Melampodium australe Loefl. 1818. Centrospermum xanthioides Kunth 1820. Acanthospermum brasilum Schrank 1836. A. xanthioides (Kunth) DC. 1891. A. australe (Loefl.) Kuntze	Southern starburr, Paraguayan starbur, Paraguay-starbur, Paraguay-bur, Sheepbur, Spiny-bur	Introduced from South America, Impersistent, Not naturalized			NYFA-Xm USDA-XX ARS Images, wsp
Acanthospermum hispidum DC. 1836. A. hispidum DC. auct. A. humile non (Sw.) DC.	Hispid starburr, Hispid starbur, Bristly starbur	Introduced from South America, Impersistent, Not naturalized			NYFA-Xm USDA-XX ARS ITIS FNA Images, wsp
Acanthospermum (excluded taxa)	Starbur	N.Y. Status	Images	Distribution	NY NPT
Acanthospermum humile (Sw.) DC. 1788. Melampodium humile Sw. 1832. Centrospermum humile (Sw.) Less. 1836. A. humile (Sw.) DC.	Low starburr, Low starbur	Introduced, N.Y. excluded, Misidentified^[1]			NYFA-XCLD USDA-X0 ARS ITIS FNA Images, wsp
↑ Specimens from New York City, identified as Acanthospermum humile, were later re-itentified as Acanthospermum hispidum.

Tribe Madieae

Known as the tarweed tribe, Madieae includes plants known as tarweeds from the western United States.

Subtribe Arnicinae

Arnica

Arnica lanceolata ssp. prima
clasping arnica

Asterales — Asteraceae — Asteroideae — Madieae — Arnicinae — Arnica
Arnica	Arnica	N.Y. Status	Images	Distribution	NY NPT
Arnica lanceolata Nutt. ssp. lanceolata Arnica mollis var. petiolaris	Lance-leaved arnica, Lanceleaf arnica, New England arnica, Hairy arnica Arnica à aigrettes brunes, Arnica soyeux	Native, Endangered			NYFA-1 USDA-NN NPT ARS BNA Images, wsp

Subtribe Madiinae

Madia

Asterales — Asteraceae — Asteroideae — Madieae — Madiinae — Madia
Madia	Tarweed	N.Y. Status	Images	Distribution	NY NPT
Madia glomerata Hook.	Mountain tarweed	Introduced, Impersistent, N. America native			NYFA-Xm USDA-NN Images, wsp
Madia sativa Molina	Chilean tarweed, Coast tarweed	Introduced, Impersistent, US West native			NYFA-Xm USDA-NX Images, wsp

Centromadia

Centromadia pungens ssp. pungens - common spikeweed

Asterales — Asteraceae — Asteroideae — Madieae — Madiinae — Centromadia
Centromadia	Spikeweed	N.Y. Status	Images	Distribution	NY NPT
Centromadia pungens (Hook. & Arn.) Greene ssp. pungens 1841. Hartmannia pungens Hook. & Arn. 1843. Hemizonia pungens (Hook. & Arn.) Torr. & A. Gray 1894. Centromadia pungens (Hook. & Arn.) Greene	Common spikeweed, Common tarweed, Pungent false tarplant	Introduced from southwestern USA, Impersistent, Not naturalized			NYFA-X USDA-N0 ARS Images, wsp

Tribe Eupatorieae

Subtribe Ageratinae

Ageratum

Ageratum houstonianum
bluemink or flossflower

Asterales — Asteraceae — Asteroideae — Eupatorieae — Ageratinae — Ageratum
Ageratum	Whiteweed	N.Y. Status	Images	Distribution	NY NPT
Ageratum houstonianum Mill. Ageratum houstonianum Mill.	Bluemink Houston's whiteweed Flossflower	Introduced, Not naturalized			NYFA-Xm USDA-X0 NPT Images, wsp
Ageratum (excluded taxa)	Whiteweed	N.Y. Status	Images	Distribution	NY NPT
Ageratum conyzoides L. Ageratum conyzoides	Goatweed Tropical whiteweed Yerba de Cabrio	Not native, N.Y. excluded, iMapInvasives			NYFA-XCLD USDA-X0 NPT Images, wsp

Subtribe Oxylobinae

Ageratina

Ageratina altissima
white snakeroot

Asterales — Asteraceae — Asteroideae — Eupatorieae — Oxylobinae — Ageratina
Ageratina	Snakeroot	N.Y. Status	Images	Distribution	NY NPT
Ageratina altissima (L.) R.M.King & H.Rob. 1753. Ageratum altissimum L. 1779. Eupatorium rugosum Houtt. 1970. Ageratina altissima (L.) R.M.King & H.Rob.	White snakeroot, Richweed Eupatoire rugueuse, Eupatoire à feuilles molles	Native, Secure			NYFA-5 USDA-NN NPT Can ARS ITIS BNA Images, wsp
Ageratina aromatica (L.) Spach	Small white snakeroot, Small-leaved white snakeroot, Lesser snakeroot	Native, Endangered, Protected (a)			NYFA-1 Images, wsp

Subtribe Mikaniinae

Mikania

Mikania scandens
climbing hempweed

Asterales — Asteraceae — Asteroideae — Eupatorieae — Mikaniinae — Mikania
Mikania	Hempvine	N.Y. Status	Images	Distribution	NY NPT
Mikania scandens (L.) Willd.	Climbing hempweed	Native, C:6, Likely secure OBL Perennial, Herb-vine			NYFA-4 USDA-NX NPT Images, wsp

Subtribe Gyptidinae

Conoclinium

Asterales — Asteraceae — Asteroideae — Eupatorieae — Gyptidinae — Conoclinium
Conoclinium	Thoroughwort	N.Y. Status	Images	Distribution	NY NPT
Conoclinium coelestinum (L.) DC. 1753. Eupatorium coelestinum L. 1836. Conoclinium coelestinum (L.) DC. 1878. Conoclinium dichotomum Chapm.	Blue mistflower, Blue boneset	Introduced, N. America native			NYFA-X USDA-NX? NPT Can ARS ITIS NSE BNA Images, wsp

Subtribe Liatridinae

Liatris

Asterales — Asteraceae — Asteroideae — Eupatorieae — Liatridinae — Liatris
Liatris	Blazing star	N.Y. Status	Images	Distribution	NY NPT
Liatris spicata (L.) Willd. var. spicata	Button blazing star, Marsh blazing star, Dense blazing star, Dense gayfeather	N. America native, Unk. NY nativity FAC Perennial, Herb-forb, Sun			NYFA-UN USDA-NN NPT NSE BNA LBJ Images, wsp
Liatris scariosa (L.) Willd. var. novae-angliae Lunell	New England blazing star	Native, C:9, Threatened			NYFA-2 USDA-N0 NPT Images, wsp
Liatris scariosa (L.) Willd. var. nieuwlandii (Lunell) E.G. Voss	Nieuwland's blazing star	Native, C:8, Endangered			NYFA-1 USDA-N0 Images, wsp
Liatris cylindracea Michx.	Slender blazing-star	Native, C:9, Endangered			NYFA-1 USDA-NN Images, wsp
Liatris aspera Michx.	Rough gayfeather, Tall blazing star, Tall gayfeather	Introduced, N. America native			NYFA-X USDA-NN Images, wsp
Liatris pycnostachya Michx. var. pycnostachya	Thick-spiked blazing star, Cattail gayfeather, Prairie blazing star	Introduced from US midwest, Midwest native			NYFA-X USDA-N0 NPT NSE BNA Images, wsp

Subtribe Eupatoriinae

The Eupatoriinae contains the Joe-Pye weeds and thoroughworts, both of which were formerly placed in the genus Eupatorium. However, in 1970, Robert M. King and Harold E. Robinson recommended moving the Joe-Pye weeds to a new genus Eupatoriadelphus.^[1] Subsequent molecular studies supported their recommendation for the taxon split, but it was determined that Eutrochium Raf. (1836) was an earlier validly published genus name for Joe-Pye weeds, so Eric E. Lamont transferred the Joe-pye weeds to Eutrochium in 2004.^[2]

↑ King, R. M., and H. Robinson, 1970b. "Studies in the Eupatorieae (Compositae). XXV. A new genus which they coined Eupatoriadelphus." Phytologia 19: 431–432.
↑ Eric E. Lamont (2004) "New combinations in Eutrochium (Asteraceae: Eupatorieae), an earlier name than Eupatoriadelphus." SIDA, contributions to botany. 21: 901-902.

Eupatorium

Eupatorium perfoliatum
common boneset

Eupatorium (thoroughwort or boneset) contains the white-flowering plants of the Eupatoriinae, native to the northern hemisphere. Most Eupatorium species found in New York are native to the state, and many are listed a protected species.

Asterales — Asteraceae — Asteroideae — Eupatorieae — Eupatoriinae — Eupatorium
Eupatorium	Thoroughwort	N.Y. Status	Images	Distribution	NY NPT
Eupatorium perfoliatum L. 1753. Eupatorium perfoliatum L. 1903. Eupatorium chapmanii Small	Common boneset, Thoroughwort, Boneset thoroughwort, Thoroughwax Eupatoire perfoliée	Native, C:3, Secure FACW Perennial, Herb-forb			NYFA-5 USDA-NN NPT ARS Trop. Images, wsp
Eupatorium hyssopifolium	Hyssop-leaved thoroughwort Hyssopleaf thoroughwort	Native, C:3, Vulnerable			NYFA-3-4 USDA-N0 Images, wsp
Eupatorium pilosum Eupatorium rotundifolium var. saundersii	Vervain thoroughwort, Rough boneset	Native, Vulnerable			NYFA-3-4 USDA-N0 Images, wsp
Eupatorium torreyanum	Torrey’s thoroughwort	Native, Threatened			NYFA-2 USDA-N0 Images, wsp
Eupatorium subvenosum	White thoroughwort	Native, Threatened			NYFA-2-3 USDA-N NPT Images, wsp
Eupatorium leucolepis	White-bracted thoroughwort, Justiceweed	Native, Endangered			NYFA-1 USDA-N0 Images, wsp
Eupatorium rotundifolium var. rotundifolium Eupatorium verbenifolium	Round-leaved boneset, Roundleaf thoroughwort	Native, Endangered			NYFA-Z USDA-N0 Images, wsp
Eupatorium pubescens Muhl. ex Willd. Eupatorium pubescens^[1] E. rotundifolium ssp. ovatum E. rotundifolium var. ovatum	Hairy boneset, Hairy thoroughwort, Roundleaf thoroughwort	Native, Endangered			NYFA-1 USDA- FNA Images, wsp
Eupatorium serotinum	Late-flowering thorough-wort, Late boneset	Native, Endangered			NYFA-2 USDA-NX Images, wsp
Eupatorium resinosum	Pine barren boneset	Native, Extirpated			NYFA-Z Images, wsp
Eupatorium sessilifolium L.	Upland boneset, Sessile-leaved boneset	Native, C:7, Unranked			NYFA-U USDA-N0 Images, wsp
Eupatorium altissimum	Tall boneset, Tall thoroughwort	N. America native, Unk. NY nativity			NYFA-X USDA-NN NPT Images, wsp
Eupatorium capillifolium (Lam.) Small ex Porter & Britton Eupatorium capillifolium (Lam.) Small ex Porter & Britton	Dog fennel Dogfennel	N. America native, Unk. NY nativity			NYFA-U USDA-N0 Images, wsp
Eupatorium cannabinum 1753. Eupatorium cannabinum L.	Hemp-agrimony Eupatoire chanvrine, Eupatoire à feuilles de chanvre	Introduced from Eurasia, northern Africa, Impersistent, Not naturalized			NYFA-X USDA-XX NEW-0 Can ARS BNA Images, wsp
Eupatorium (excluded taxa)	Thoroughwort	N.Y. Status	Images	Distribution	NY NPT
Eupatorium album L. Eupatorium album L. Eupatorium glandulosum Michx. Eupatorium fernaldii Godfrey Eupatorium petaloideum	White thoroughwort	Native, Endangered			NYFA-XCLD USDA-N NPT ITIS Images, wsp Note: CA
↑ The New York Flora Atlas lists E. rotundifolium var. ovatum as E. pubescens. The Flora of North America says "Variety ovatum is an apomictic polyploid that may be a derivative of Eupatorium rotundifolium and E. sessilifolium. Morphologically, it is closer to E. rotundifolium than to E. sessilifolium; the name E. pubescens has priority at species rank."

Eutrochium

Eutrochium maculatum
spotted Joe-Pye weed

Eutrochium (commonly called Joe Pye weed) contains the purple-flowered North American species of the Eupatoriinae. Until 2004, these species were included in the genus Eupatorium and later Eupatoriadelphus.

Asterales — Asteraceae — Asteroideae — Eupatorieae — Eupatoriinae — Eutrochium
Eutrochium	Joe-Pye weed	N.Y. Status	Images	Distribution	NY NPT
Eutrochium maculatum (L.) E.E.Lamont var. maculatum 1755. Eupatorium maculatum L. 1970. Eupatoriadelphus maculatus (L.) R.M.King & H.Rob. 2004. Eutrochium maculatum (L.) E.E.Lamont	Spotted Joe Pye weed Spotted joe pye weed Spotted Joe-Pye weed Eupatoire maculée	Native, C:4, Secure OBL Perennial, Herb-forb, Part shade			NYFA-5 USDA-N NPT ARS ITIS Trop. NSE BNA LBJ Images, wsp
Eutrochium maculatum (L.) E.E.Lamont var. foliosum 1908. Eupatorium purpureum var. foliosum Fernald 1920. Eupatorium maculatum var. foliosum (Fernald) Wiegand 2004. Eutrochium maculatum var. foliosum (Fernald) E.E. Lamont	Leafy Joe Pye weed Spotted joe pye weed Eupatoire feuillue	Native, C:4, Unranked OBL Perennial, Herb-forb			NYFA-Nu USDA-NN ARS NSE BNA Images, wsp
Eutrochium purpureum (L.) E.E.Lamont var. purpureum 1753. Eupatorium purpureum L. 1917. Cunigunda purpurea (L.) Lunell 1970. Eupatoriadelphus purpureus (L.) R.M.King & H.Rob. 2004. Eutrochium purpureum (L.) E.E.Lamont	Purple Joe-Pye weed, Sweet-scented Joe-Pye weed, Sweetscented joe pye weed, Green-stem joe-pyeweed	Native, C:7, Secure FAC Perennial, Herb-forb			NYFA-5 USDA-NN NPT ARS FNA Trop. NSE BNA Images, wsp
Eutrochium fistulosum (Barratt) E.E.Lamont 1841. Eupatorium purpureum var. angustifolium Torr. & A.Gray 1849. Eupatorium fistulosum Barratt 1970. Eupatoriadelphus fistulosus (Barratt) R.M.King & H.Rob. 2004. Eutrochium fistulosum (Barratt) E.E.Lamont	Hollow Joe-Pye weed, Trumpetweed	Native, C:1, Secure FACW Perennial, Herb-forb			NYFA-5 USDA-NN NPT ARS FNA Trop. NSE BNA Images, wsp
Eutrochium dubium (Willd. ex Poir.) E.E.Lamont 1812. Eupatorium dubium Willd. ex Poir. 1970. Eupatoriadelphus dubius (Willd. ex Poir.) R.M.King & H.Rob. 2004. Eutrochium dubium (Willd. ex Poir.) E.E.Lamont	Coastal plain Joe-Pye weed, Joe-Pye thoroughwort	Native, C:10, Vulnerable FACW Perennial, Herb-forb			NYFA-3 USDA-N NPT ARS FNA Trop. NSE BNA Images, wsp
Eutrochium (excluded taxa)	Joe-Pye weed	N.Y. Status	Images	Distribution	NY NPT
Eutrochium maculatum (L.) E.E.Lamont var. bruneri Eupatoriadelphus maculatus var. bruneri Eupatorium bruneri A.Gray	Spotted Joe-Pye weed	N. America native, N.Y. excluded			USDA-NN NSE BNA Images, wsp

Eutrochium identification

Eutrochium Joe-Pye weed	Floret color	Florets per head	Capitu- lescence shape	Stem color (besides green)	Stem glaucousness (white bloom)	Central stem cavity (hollow/solid)	Leaf taper to petiole	Leaf main veins	Max stem height
E. purpureum (purple)	pale pink, purplish	(4-)5 - 8(-10)	round, domed	dark purple at nodes	not/scarcely glaucous	solid or thin cavity at base	gradual taper	1 main vein	2 m
E. fistulosum (hollow)	pink- purple	(4-)5 - 8(-10)	round, domed	purple throughout	prominently glaucous	large cavity (hollow)	gradual taper	1 main vein	3 m
E. maculatum (spotted)	purplish	(8-)9 - 20(-22)	flatish top	purple spots or all purple	rarely glaucous	solid or thin cavity at base	gradual taper	1 main vein	2 m
E. dubium (coastal-plain)	purple	(4-)5 - 8(-10)	round, domed	fine purple spots/streaks			abruptly narrowed	3 main veins	1 m

← Coreopsideae … Ambrosieae	Flora of New York — Millerieae, Madieae, Eupatorieae				→ Apiales
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[1] Specimens from New York City, identified as Acanthospermum humile, were later re-itentified as Acanthospermum hispidum.